From Wikipedia, the free encyclopedia
The identity of the longest word in English depends on the definition of a word and of length.
Words may be derived naturally from the language’s roots or formed by coinage and construction. Additionally, comparisons are complicated because place names may be considered words, technical terms may be arbitrarily long, and the addition of suffixes and prefixes may extend the length of words to create grammatically correct but unused or novel words.
The length of a word may also be understood in multiple ways. Most commonly, length is based on orthography (conventional spelling rules) and counting the number of written letters. Alternate, but less common, approaches include phonology (the spoken language) and the number of phonemes (sounds).
Word | Letters | Meaning | Claim | Dispute |
---|---|---|---|---|
methionylthreonylthreonylglutaminylalanyl…isoleucine | 189,819 | The chemical composition of titin, the largest known protein | Longest known word overall by magnitudes. Attempts to say the entire word have taken two[1] to three and a half hours.[2] | Technical; not in dictionary; whether this should actually be considered a word is disputed |
methionylglutaminylarginyltyrosylglutamyl…serine | 1,909 | The chemical name of E. coli TrpA (P0A877) | Longest published word[3] | Technical |
lopadotemachoselachogaleokranioleipsano…pterygon | 183 | A fictional dish of food | Longest word coined by a major author,[4] the longest word ever to appear in literature[5] | Contrived nonce word; not in dictionary; Ancient Greek transliteration |
pneumonoultramicroscopicsilicovolcanoconiosis | 45 | The disease silicosis | Longest word in a major dictionary[6] | Contrived coinage to make it the longest word; technical, but only mentioned and never actually used in communication |
supercalifragilisticexpialidocious | 34 | Unclear in source work, has been cited as a nonsense word | Made popular in the Mary Poppins film and musical[7] | Contrived coinage |
pseudopseudohypoparathyroidism | 30 | A hereditary medical disorder | Longest non-contrived word in a major dictionary[8] | Technical |
antidisestablishmentarianism | 28 | The political position of opposing disestablishment | Longest non-contrived and nontechnical word[9] | Not all dictionaries accept it due to lack of usage.[10] |
honorificabilitudinitatibus | 27 | The state of being able to achieve honors | Longest word in Shakespeare’s works; longest word in the English language featuring alternating consonants and vowels[11] | Latin |
Major dictionaries
The longest word in any of the major English language dictionaries is pneumonoultramicroscopicsilicovolcanoconiosis (45 letters), a word that refers to a lung disease contracted from the inhalation of very fine silica particles,[12] specifically from a volcano; medically, it is the same as silicosis. The word was deliberately coined to be the longest word in English, and has since been used[citation needed] in a close approximation of its originally intended meaning, lending at least some degree of validity to its claim.[6]
The Oxford English Dictionary contains pseudopseudohypoparathyroidism (30 letters).
Merriam-Webster’s Collegiate Dictionary does not contain antidisestablishmentarianism (28 letters), as the editors found no widespread, sustained usage of the word in its original meaning. The longest word in that dictionary is electroencephalographically (27 letters).[13]
The longest non-technical word in major dictionaries is floccinaucinihilipilification at 29 letters. Consisting of a series of Latin words meaning «nothing» and defined as «the act of estimating something as worthless»; its usage has been recorded as far back as 1741.[14][15][16]
Ross Eckler has noted that most of the longest English words are not likely to occur in general text, meaning non-technical present-day text seen by casual readers, in which the author did not specifically intend to use an unusually long word. According to Eckler, the longest words likely to be encountered in general text are deinstitutionalization and counterrevolutionaries, with 22 letters each.[17]
A computer study of over a million samples of normal English prose found that the longest word one is likely to encounter on an everyday basis is uncharacteristically, at 20 letters.[18]
The word internationalization is abbreviated «i18n», the embedded number representing the number of letters between the first and the last.[19][20][21]
Creations of long words
Coinages
In his play Assemblywomen (Ecclesiazousae), the ancient Greek comedic playwright Aristophanes created a word of 171 letters (183 in the transliteration below), which describes a dish by stringing together its ingredients:
Henry Carey’s farce Chrononhotonthologos (1743) holds the opening line: «Aldiborontiphoscophornio! Where left you Chrononhotonthologos?»
Thomas Love Peacock put these creations into the mouth of the phrenologist Mr. Cranium in his 1816 book Headlong Hall: osteosarchaematosplanchnochondroneuromuelous (44 characters) and osseocarnisanguineoviscericartilaginonervomedullary (51 characters).
James Joyce made up nine 100-letter words plus one 101-letter word in his novel Finnegans Wake, the most famous of which is Bababadalgharaghtakamminarronnkonnbronntonnerronntuonnthunntrovarrhounawnskawntoohoohoordenenthurnuk. Appearing on the first page, it allegedly represents the symbolic thunderclap associated with the fall of Adam and Eve. As it appears nowhere else except in reference to this passage, it is generally not accepted as a real word. Sylvia Plath made mention of it in her semi-autobiographical novel The Bell Jar, when the protagonist was reading Finnegans Wake.
«Supercalifragilisticexpialidocious», the 34-letter title of a song from the movie Mary Poppins, does appear in several dictionaries, but only as a proper noun defined in reference to the song title. The attributed meaning is «a word that you say when you don’t know what to say.» The idea and invention of the word is credited to songwriters Robert and Richard Sherman.
Agglutinative constructions
The English language permits the legitimate extension of existing words to serve new purposes by the addition of prefixes and suffixes. This is sometimes referred to as agglutinative construction. This process can create arbitrarily long words: for example, the prefixes pseudo (false, spurious) and anti (against, opposed to) can be added as many times as desired. More familiarly, the addition of numerous «great»s to a relative, such as «great-great-great-great-grandparent», can produce words of arbitrary length. In musical notation, an 8192nd note may be called a semihemidemisemihemidemisemihemidemisemiquaver.
Antidisestablishmentarianism is the longest common example of a word formed by agglutinative construction.
Technical terms
A number of scientific naming schemes can be used to generate arbitrarily long words.
The IUPAC nomenclature for organic chemical compounds is open-ended, giving rise to the 189,819-letter chemical name Methionylthreonylthreonyl…isoleucine for the protein also known as titin, which is involved in striated muscle formation. In nature, DNA molecules can be much bigger than protein molecules and therefore potentially be referred to with much longer chemical names. For example, the wheat chromosome 3B contains almost 1 billion base pairs,[22] so the sequence of one of its strands, if written out in full like Adenilyladenilylguanilylcystidylthymidyl…, would be about 8 billion letters long. The longest published word, Acetylseryltyrosylseryliso…serine, referring to the coat protein of a certain strain of tobacco mosaic virus (P03575), is 1,185 letters long, and appeared in the American Chemical Society’s Chemical Abstracts Service in 1964 and 1966.[23] In 1965, the Chemical Abstracts Service overhauled its naming system and started discouraging excessively long names. In 2011, a dictionary broke this record with a 1909-letter word describing the trpA protein (P0A877).[3]
John Horton Conway and Landon Curt Noll developed an open-ended system for naming powers of 10, in which one sexmilliaquingentsexagintillion, coming from the Latin name for 6560, is the name for 103(6560+1) = 1019683. Under the long number scale, it would be 106(6560) = 1039360.
Gammaracanthuskytodermogammarus loricatobaicalensis is sometimes cited as the longest binomial name—it is a kind of amphipod. However, this name, proposed by B. Dybowski, was invalidated by the International Code of Zoological Nomenclature in 1929 after being petitioned by Mary J. Rathbun to take up the case.[24]
Myxococcus llanfairpwllgwyngyllgogerychwyrndrobwllllantysiliogogogochensis is the longest accepted binomial name for an organism. It is a bacterium found in soil collected at Llanfairpwllgwyngyll (discussed below). Parastratiosphecomyia stratiosphecomyioides is the longest accepted binomial name for any animal, or any organism visible with the naked eye. It is a species of soldier fly.[25] The genus name Parapropalaehoplophorus (a fossil glyptodont, an extinct family of mammals related to armadillos) is two letters longer, but does not contain a similarly long species name.
Aequeosalinocalcalinoceraceoaluminosocupreovitriolic, at 52 letters, describing the spa waters at Bath, England, is attributed to Dr. Edward Strother (1675–1737).[26] The word is composed of the following elements:
- Aequeo: equal (Latin, aequo[27])
- Salino: containing salt (Latin, salinus)
- Calcalino: calcium (Latin, calx)
- Ceraceo: waxy (Latin, cera)
- Aluminoso: alumina (Latin)
- Cupreo: from «copper»
- Vitriolic: resembling vitriol
Notable long words
Place names
The longest officially recognized place name in an English-speaking country is Taumatawhakatangihangakoauauotamateaturipukakapikimaungahoronukupokaiwhenuakitanatahu (85 letters), which is a hill in New Zealand. The name is in the Māori language. A widely recognized version of the name is Taumatawhakatangihangakoauauotamateaturipukakapikimaungahoronukupokaiwhenuakitanatahu (85 letters), which appears on the signpost at the location (see the photo on this page). In Māori, the digraphs ng and wh are each treated as single letters.
In Canada, the longest place name is Dysart, Dudley, Harcourt, Guilford, Harburn, Bruton, Havelock, Eyre and Clyde, a township in Ontario, at 61 letters or 68 non-space characters.[28]
The 58-letter name Llanfairpwllgwyngyllgogerychwyrndrobwllllantysiliogogogoch is the name of a town on Anglesey, an island of Wales. In terms of the traditional Welsh alphabet, the name is only 51 letters long, as certain digraphs in Welsh are considered as single letters, for instance ll, ng and ch. It is generally agreed, however, that this invented name, adopted in the mid-19th century, was contrived solely to be the longest name of any town in Britain. The official name of the place is Llanfairpwllgwyngyll, commonly abbreviated to Llanfairpwll or Llanfair PG.
The longest non-contrived place name in the United Kingdom which is a single non-hyphenated word is Cottonshopeburnfoot (19 letters) and the longest which is hyphenated is Sutton-under-Whitestonecliffe (29 characters).
The longest place name in the United States (45 letters) is Chargoggagoggmanchauggagoggchaubunagungamaugg, a lake in Webster, Massachusetts. It means «Fishing Place at the Boundaries – Neutral Meeting Grounds» and is sometimes facetiously translated as «you fish your side of the water, I fish my side of the water, nobody fishes the middle». The lake is also known as Webster Lake.[29] The longest hyphenated names in the U.S. are Winchester-on-the-Severn, a town in Maryland, and Washington-on-the-Brazos, a notable place in Texas history. The longest single-word town names in the U.S. are Kleinfeltersville, Pennsylvania and Mooselookmeguntic, Maine.
The longest official geographical name in Australia is Mamungkukumpurangkuntjunya.[30] It has 26 letters and is a Pitjantjatjara word meaning «where the Devil urinates».[31]
Liechtenstein is the longest country name with single name in English. The second longest country name with single name in English is Turkmenistan. There are longer country names if one includes ones with spaces.
Personal names
Guinness World Records formerly contained a category for longest personal name used.
- From about 1975 to 1985, the recordholder was Adolph Blaine Charles David Earl Frederick Gerald Hubert Irvin John Kenneth Lloyd Martin Nero Oliver Paul Quincy Randolph Sherman Thomas Uncas Victor William Xerxes Yancy Zeus Wolfeschlegelsteinhausenbergerdorffvoralternwarengewissenhaftschaferswessenschafewarenwohlgepflegeundsorgfaltigkeitbeschutzenvonangreifendurchihrraubgierigfeindewelchevoralternzwolftausendjahresvorandieerscheinenwanderersteerdemenschderraumschiffgebrauchlichtalsseinursprungvonkraftgestartseinlangefahrthinzwischensternartigraumaufdersuchenachdiesternwelchegehabtbewohnbarplanetenkreisedrehensichundwohinderneurassevonverstandigmenschlichkeitkonntefortplanzenundsicherfreuenanlebenslanglichfreudeundruhemitnichteinfurchtvorangreifenvonandererintelligentgeschopfsvonhinzwischensternartigraum, Senior (746 letters), also known as Wolfe+585, Senior.
- After 1985 Guinness briefly awarded the record to a newborn girl with a longer name. The category was removed shortly afterward.
Long birth names are often coined in protest of naming laws or for other personal reasons.
- The naming law in Sweden was challenged by parents Lasse Diding and Elisabeth Hallin, who proposed the given name «Brfxxccxxmnpcccclllmmnprxvclmnckssqlbb11116» for their child (pronounced [ˈǎlːbɪn], 43 characters), which was rejected by a district court in Halmstad, southern Sweden.
Words with certain characteristics of notable length
- Schmaltzed and strengthed (10 letters) appear to be the longest monosyllabic words recorded in The Oxford English Dictionary, while scraunched and scroonched appear to be the longest monosyllabic words recorded in Webster’s Third New International Dictionary; but squirrelled (11 letters) is the longest if pronounced as one syllable only (as permitted in The Shorter Oxford English Dictionary and Merriam-Webster Online Dictionary at squirrel, and in Longman Pronunciation Dictionary). Schtroumpfed (12 letters) was coined by Umberto Eco, while broughammed (11 letters) was coined by William Harmon after broughamed (10 letters) was coined by George Bernard Shaw.
- Strengths is the longest word in the English language containing only one vowel letter.[32]
- Euouae, a medieval musical term, is the longest English word consisting only of vowels, and the word with the most consecutive vowels. However, the «word» itself is simply a mnemonic consisting of the vowels to be sung in the phrase «seculorum Amen» at the end of the lesser doxology. (Although u was often used interchangeably with v, and the variant «Evovae» is occasionally used, the v in these cases would still be a vowel.)
- The longest words with no repeated letters are dermatoglyphics and uncopyrightable.[33]
- The longest word whose letters are in alphabetical order is the eight-letter Aegilops, a grass genus. However, this is arguably a proper noun. There are several six-letter English words with their letters in alphabetical order, including abhors, almost, begins, biopsy, chimps and chintz.[34] There are few 7-letter words, such as «billowy» and «beefily». The longest words whose letters are in reverse alphabetical order are sponged, wronged and trollied.
- The longest words recorded in OED with each vowel only once, and in order, are abstemiously, affectiously, and tragediously (OED). Fracedinously and gravedinously (constructed from adjectives in OED) have thirteen letters; Gadspreciously, constructed from Gadsprecious (in OED), has fourteen letters. Facetiously is among the few other words directly attested in OED with single occurrences of all six vowels (counting y as a vowel).
- The longest single palindromic word in English is rotavator, another name for a rotary tiller for breaking and aerating soil.
Typed words
- The longest words typable with only the left hand using conventional hand placement on a QWERTY keyboard are tesseradecades, aftercataracts, dereverberated, dereverberates[35] and the more common but sometimes hyphenated sweaterdresses.[34] Using the right hand alone, the longest word that can be typed is johnny-jump-up, or, excluding hyphens, monimolimnion[36] and phyllophyllin.
- The longest English word typable using only the top row of letters has 11 letters: rupturewort. The word teetertotter (used in North American English) is longer at 12 letters, although it is usually spelled with a hyphen.
- The longest using only the middle row is shakalshas (10 letters). Nine-letter words include flagfalls; eight-letter words include galahads and alfalfas.
- Since the bottom row contains no vowels, no standard words can be formed. [37]
- The longest words typable by alternating left and right hands are antiskepticism and leucocytozoans respectively.[34]
- On a Dvorak keyboard, the longest «left-handed» words are epopoeia, jipijapa, peekapoo, and quiaquia.[38] Other such long words are papaya, Kikuyu, opaque, and upkeep.[39] Kikuyu is typed entirely with the index finger, and so the longest one-fingered word on the Dvorak keyboard. There are no vowels on the right-hand side, and so the longest «right-handed» word is crwths.
See also
- Lipogram
- List of long species names
- List of the longest English words with one syllable
- Longest English sentence
- Longest word in French
- Longest word in Romanian
- Longest word in Spanish
- Longest word in Turkish
- Number of words in English
- Scriptio continua
- Sesquipedalianism
- Donaudampfschiffahrtselektrizitätenhauptbetriebswerkbauunterbeamtengesellschaft, longest published word in German
References
- ^ «Reading The Longest English Word (190,000 Characters)». YouTube. Archived from the original on 2021-11-10. Retrieved 2 August 2020.
- ^ «World’s longest word takes 3.5 hours to pronounce». CW39 Houston. 2012-12-08. Retrieved 2020-05-18.
- ^ a b Colista Moore (2011). Student’s Dictionary. p. 524. ISBN 978-1-934669-21-1.
- ^ see separate article Lopado…pterygon
- ^ Donald McFarlan; Norris Dewar McWhirter; David A. Boeh (1989). Guinness book of world records: 1990. Sterling. p. 129. ISBN 978-0-8069-5790-6.
- ^ a b Coined around 1935 to be the longest word; press reports on puzzle league members legitimized it somewhat. First appeared in the MWNID supplement, 1939. Today OED and several others list it, but citations are almost always as «longest word». More detail at pneumonoultramicroscopicsilicovolcanoconiosis.
- ^ «Merriam Webster: Supercalifragilisticexpialidocious».
- ^ «What is the longest English word?». AskOxford. Archived from the original on 2008-10-22. Retrieved 2010-08-22.
- ^ «What is the longest English word?». oxforddictionaries.com.[dead link]
- ^ «Merriam Webster: «Antidisestablishmentarianism is not in the dictionary.»«.
- ^ «Cool, Strange, and Interesting Facts,» fact 99. InnocentEnglish.com. Retrieved 2019-03-13.
- ^ «pneumonoultramicroscopicsilicovolcanoconiosis – definition of pneumonoultramicroscopicsilicovolcanoconiosis in English from the Oxford dictionary». oxforddictionaries.com. Archived from the original on 2012-07-19.
- ^ «The Longest Word in the Dictionary» (Video). Ask the Editor. Merriam-Webster. Archived from the original on 21 November 2013. Retrieved 14 November 2013.
- ^ «Floccinaucinihilipilification» by Michael Quinion World Wide Words Archived 2006-08-21 at the Wayback Machine;
- ^ The Guinness Book of Records, in its 1992 and previous editions, declared the longest real word in the English language to be floccinaucinihilipilification. More recent editions of the book have acknowledged pneumonoultramicroscopicsilicovolcanoconiosis. What is the longest English word? — Oxford Dictionaries Online Archived 2006-08-26 at the Wayback Machine
- ^ In recent times its usage has been recorded in the proceedings of the United States Senate by Senator Robert Byrd Discussion between Sen. Moynihan and Sen. Byrd «Mr. President, may I say to the distinguished Senator from New York, I used that word on the Senate floor myself 2 or 3 years ago. I cannot remember just when or what the occasion was, but I used it on that occasion to indicate that whatever it was I was discussing it was something like a mere trifle or nothing really being of moment.» Congressional Record June 17, 1991, p. S7887, and at the White House by Bill Clinton’s press secretary Mike McCurry, albeit sarcastically. December 6, 1995, White House Press Briefing in discussing Congressional Budget Office estimates and assumptions: «But if you – as a practical matter of estimating the economy, the difference is not great. There’s a little bit of floccinaucinihilipilification going on here.»
- ^ Eckler, R. Making the Alphabet Dance, p 252, 1996.
- ^ «Longest Common Words – Modern». Maltron.com. Archived from the original on 27 April 2009. Retrieved 2010-08-22.
- ^ «Glossary of W3C Jargon». World Wide Web Consortium. Archived from the original on 2008-10-25. Retrieved 2008-10-13.
- ^ «Origin of the Abbreviation I18n». Archived from the original on 2014-06-27.
- ^ «Localization vs. Internationalization». World Wide Web Consortium. Archived from the original on 2016-04-03.
- ^ Paux et al. (2008) Science, Vol. 322 (5898) 101-104. A Physical Map of the 1-Gigabase Bread Wheat Chromosome 3B Paux, Etienne; Sourdille, Pierre; Salse, Jérôme; Saintenac, Cyrille; Choulet, Frédéric; Leroy, Philippe; Korol, Abraham; Michalak, Monika; Kianian, Shahryar; Spielmeyer, Wolfgang; Lagudah, Evans; Somers, Daryl; Kilian, Andrzej; Alaux, Michael; Vautrin, Sonia; Bergès, Hélène; Eversole, Kellye; Appels, Rudi; Safar, Jan; Simkova, Hana; Dolezel, Jaroslav; Bernard, Michel; Feuillet, Catherine (2008). «A Physical Map of the 1-Gigabase Bread Wheat Chromosome 3B». Science. 322 (5898): 101–104. Bibcode:2008Sci…322..101P. doi:10.1126/science.1161847. PMID 18832645. S2CID 27686615. Archived from the original on 2015-09-03. Retrieved 2012-12-01.
- ^ Chemical Abstracts Formula Index, Jan.-June 1964, Page 967F; Chemical Abstracts 7th Coll. Formulas, C23H32-Z, 56-65, 1962–1966, Page 6717F
- ^ «Opinion 105. Dybowski’s (1926) Names of Crustacea Suppressed». Opinions Rendered by the International Commission on Zoological Nomenclature: Opinions 105 to 114. Smithsonian Miscellaneous Collections. Vol. 73. 1929. pp. 1–3. hdl:10088/23619. BHL page 8911139.
- ^ rjk. «World’s longest name of an animal. Parastratiosphecomyia stratiosphecomyioides Stratiomyid Fly Soldier Fly». thelongestlistofthelongeststuffatthelongestdomainnameatlonglast.com. Archived from the original on 2011-11-17. Retrieved 2011-12-17.
- ^ cited in some editions of the Guinness Book of Records as the longest word in English, see Askoxford.com on the longest English word
- ^ [1][dead link]
- ^ «GeoNames Government of Canada site». Archived from the original on 2009-02-06.
- ^ Belluck, Pam (2004-11-20). «What’s the Name of That Lake? It’s Hard to Say». The New York Times.
- ^ «Geoscience Australia Gazetteer». Archived from the original on 2007-10-01.
- ^ «South Australian State Gazetteer». Archived from the original on 2007-10-01.
- ^ «Guinness Records».
- ^ «Longest Word Without Repeating Letters». December 2014.
- ^ a b c «Typewriter Words». Questrel.com. Archived from the original on 2010-09-27. Retrieved 2010-08-22.
- ^ «Science Links Japan | Two Unique Aftercataracts Requiring Surgical Removal». Sciencelinks.jp. 2009-03-18. Archived from the original on 2011-02-17. Retrieved 2010-08-22.
- ^ «Dictionary entry for monimolimnion, a word that, at 13 letters, is longer than any of the words linked in the source above». Archived from the original on 2009-09-09. Retrieved 2009-08-15.
- ^ «Word Records». Fun-with-words.com. Archived from the original on 2012-08-26. Retrieved 2012-08-13.
- ^ «Typewriter Words». Wordnik.com. Archived from the original on 2011-07-17. Retrieved 2011-01-15.
- ^ «The Dvorak Keyboard and You». Theworldofstuff.com. Archived from the original on 2010-08-20. Retrieved 2010-08-22.
External links
This audio file was created from a revision of this article dated 8 January 2011, and does not reflect subsequent edits.
- A Collection of Word Oddities and Trivia – Long words
- Long words (chemical names)
- Long words (place names)
- What is the longest English word?, AskOxford.com «Ask the Experts»
- What is the Longest Word?, Fun-With-Words.com
- Full chemical name of titin.
- Taxonomy of Wordplay
Ready to take your Scrabble skills to the next level? This list of the longest words in the English language could score you major points on your next game — if you can remember how to spell them.
Some of the words that qualify for the title take hours to pronounce, like the 189,819-letter word for the protein Titin. Additionally, many of the longest words are medical terms, so we have excluded some of them to allow for more variety. The end result is a list of fascinatingly lengthy words that will make your vocabulary downright sesquipedalian.
Antidisestablishmentarianism
Part of Speech: noun
Definition: opposition to the disestablishment of the Church of England
Origins: While the word originated in 19th century Britain, it is now used to refer to any opposition to a government withdrawing support from a religious organization. Though rarely used in casual conversation, the word was featured in the Duke Ellington song, “You’re Just an Old Antidisestablishmentarianist.”
Floccinaucinihilipilification
Part of Speech: noun
Definition: the act of defining or estimating something as worthless
Origins: This word stems from the combination of four Latin words, all of which signify that something has little value: flocci, nauci, nihili, pilifi. This style of word creation was popular in Britain in the 1700s.
Pneumonoultramicroscopicsilicovolcanoconiosis
Part of Speech: noun
Definition: an invented word said to mean a lung disease caused by inhaling a fine dust
Origins: This word emerged in the late 1930s, and was said to be invented by Everett K. Smith, president of the National Puzzlers’ League, in an imitation of very long medical terms. It is not found in real medical usage.
Pseudopseudohypoparathyroidism
Part of Speech: noun
Definition: an inherited disorder similar to pseudohypoparathyroidism
Origins: This genetic disorder causes “short stature, round face and short hand bones,” according to the National Institutes of Health. Despite having a similar name, it is not the same as pseudohypoparathyroidism.
Psychoneuroendocrinological
Part of Speech: adjective
Definition: of or related to to the branch of science concerned with the relationships between psychology, the nervous system, and the endocrine system
Origins: This term was first seen in the 1970s in Journal of Neurological Science, a medical journal.
Sesquipedalian
Part of Speech: adjective
Definition: having many syllables or characterized by the use of long words
Origins: The Roman poet Horace used this term to caution young poets against relying on words that used a large number of letters. It was adopted in the 17th century by poets to ridicule their peers who used lengthy words.
Hippopotomonstrosesquippedaliophobia
Part of Speech: noun
Definition: fear of long words
Fun Fact: This word is most often used in humorous contexts. It is an extension of the word sesquipedalophobia, which has the same meaning and is more often used in a formal context.
Incomprehensibilities
Part of Speech: noun
Definition: things that are impossible to understand or comprehend
Fun Fact: In the 1990s, this word was named the longest word in common usage.
Uncopyrightable
Part of Speech: adjective
Definition: not able or allowed to be protected by copyright
Fun Fact: This word is one of the longest isograms (a word that does not repeat letters) in the English language.
Dermatoglyphics
Part of Speech: noun
Definition: the scientific study of hands, including fingerprints, lines, mounts, and shapes
Fun Fact: Unlike palmistry, this study is based in science and is often used in criminology as a way to identify both perpetrators and victims.
Euouae
Part of Speech: noun
Definition: a type of cadence in medieval music
Fun Fact: While this word might not look as impressive as others on this list, it’s the longest word in the English language to be composed entirely of vowels. (It’s also the word with the longest string of vowels.)
Psychophysicotherapeutics
Part of Speech: noun
Definition: a therapeutic approach that integrates both the mind and body
Fun Fact: While the Oxford Dictionary does not provide an official definition of this word, it is included on their list of the longest words in the English language.
Otorhinolaryngological
Part of Speech: adjective
Definition: of or relating to the medical specialization involving the ear, nose, and throat
Fun Fact: This medical specialization is more commonly known by its acronym, ENT.
Прежде чем вы начнете читать статью, ответьте на вопрос: Сколько букв в самом длинном слове английского языка? 50? 75? 100? 220? В любом случае, запомните эту цифру.
Трудно поверить, но в самом коротком длинном слове в английском языке «всего» 27(!) букв. Мы начнем с самого короткого слова, а закончим самым длинным:
1. Honorificabilitudinitatibus
Это слово придумал Уильям Шекспир, и найти его можно в комедии «Бесплодные усилия любви» (Love’sLabour’s Lost). Имеет ли это слово значение? Да. Оно означает «достойный почестей, признания», и вы найдете его в словарях, хотя оно и маркируется как устаревшее.
2. Antidisestablishmentarianism
Это слово (28 букв!) вы найдете в словаре, и переводится оно следующим образом – движение за не разделение государства и церкви. Данное движение появилось в Англии в середине 1860-ых. Хотя слово и относится к устаревшим (архаичным) словам, оно встречается в научной (исторической) и художественной литературе.
3. Floccinaucinihilipilification
В этом слове 29 букв и состоит оно из корней латинских слов. Что оно означает? Вы его без труда найдете в словарях, а означает оно «опорочивание, дискредитация».
4. Pseudopseudohypoparathyroidism
В этом слове 30 букв. И обозначает оно один из типов генетического нарушения. Этот тип схож с тем, что в медицине называют псевдогипопаратиреоз, но при нем не наблюдается дефицит кальция. Не вдаваясь в медицинские детали, лучше не иметь у себя в организме то, что называется словом из 30 букв.
5. Supercalifragilisticexpialidocious
Несмотря на то, что в этом слове 34 буквы, оно весьма популярно, так как встречается в песне, которую поет Мэри Поппинс в мюзикле «Мэри Поппинс». В словарях вы найдете такой перевод — суперархиэкстраультрамегаграндиозно. Авторство принадлежит Ричарду и Роберту Шерманам, написавшим тексты песен для мюзикла. В слове угадывается некоторый оттенок учености, хотя сама героиня книги говорила, что его стоит произносить, когда совсем нечего сказать.
6. Pneumonoultramicroscopicsilicovolcanoconiosis
В этом медицинском термине 45 букв! И означает он заболевание легких, вызванное вдыханием пыли, содержащей свободную двуокись кремния. Это слово зафиксировано в словарях, но в медицинской практике пользуются очень коротким словом – silicosis (силикоз или силикатоз). Доктора, которые пишут в истории болезни такое длинное слово, очевидно, получают почасовую зарплату.
7. Aequeosalinocalcalinoceraceoaluminosocupreovitriolic
Это слово из 52 букв придумал Д-р Эдвард Строзер, для того чтобы описать состав воды для spa-процедур, которые можно получить в английском городе Бат (Bath). И сделал он это при помощи одного слова! Если расшифровать это слово, то получится – «равномерно соленая», «содержит кальций», «содержит алюминий и медь», «имеет восковый блеск». Конечно, вряд ли вы увидите это слово в научных статьях или услышите в речи, разве что в самих spa-салонах увидите его на постерах.
8. Lopadotemachoselachogaleokranioleipsanodrimhypotr
immatosilphioparaomelito-katakechymenokichlepikossyphophattoperisteralektryonoptekephalliokigklopeleiolagoio-siraiobaphetraganopterygon
В этом слове 171 буква, а авторство принадлежит древнегреческому автору Аристофану. В одной из комедий он приводит придуманное им самим блюдо. В основе – названия необычных ингредиентов: акулы, голуби, мед и всевозможные фрагменты тел животных, надо сказать, не всегда аппетитные.
В начале статьи мы попросили вас предположить, сколько букв в самом длинном слове английского языка. Какую цифру вы выбрали? Смеем предположить, что вы ошиблись. Потому что самое длинное английское слово —
9. Methionylglutaminylarginyltyrosylglutamylserylleucylphenyl—
alanylalanylglutaminylleucyllysylglutamylarginyllysylglutamyl—
glycylalanylphenylalanylvalylprolylphenylalanylvalylthreonyl—
leucylglycylaspartylprolylglycylisoleucylglutamylglutaminyl—
serylleucyllysylisoleucylaspartylthreonylleucylisoleucylglutamy—
lalanylglycylalanylaspartylalanylleucylglutamylleucylglycylisoleucyl—
prolylphenylalanylserylaspartylprolylleucylalanylaspartylglycylproly—
lthreonylisoleucylglutaminylaspfraginylalanylthreonylleucylarginy—
lalanylphenylalanylalanylalanylglycylvalylthreonylprolylalanyl—
glutaminylcysteinylphenylalanylglutamylmethionylleucylalany—
lleucylisoleucylarginylglutaminyllysylhistidylprolylthreonylisoleucyl—
prolylisoleucylglycylleucylleucylmethionyltyrosylalanylasparaginy—
lleucylvalylphenylalanylasparaginyllysylglycylisoleucylaspartyl—glutamylphenylalanyltyrosylalanylglutaminylcysteinylglutamylly—sylvalylglycylvalylaspartylserylvalylleucylvalylalanylaspartylvalyl—prolylvalylglutaminylglutamylserylalanylprolylphenylalanylarginyl—glutaminylalanylalanylleucylarginylhistidylasparaginylvalylalanyl—prolylisoleucylphenylalanylisoleucylcysteinylprolylprolylaspartylalanyl—aspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanyl—seryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylseryl—arginylalanylglycylvalylthreonylglycylalanylglutamylasparaginyl—arginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalyl—alanyllysylleucyllysylglutamyltyrosylasparaginylalanylalanylprolyl—prolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanyl—prolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanyl—glycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisoleucyl—valyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginy—lisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanyl—leucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanyl—alanylthreonylarginylserine
Да, это одно слово и в нем 1913 букв (!). Это химическое название триптофансинтетазы, белка, в составе которого 27 аминокислот.
Конечно, нет смысла составлять такие длинные слова, прежде всего из-за их непрактичности. Поэтому в языке их не так уж много. Но на досуге можно попробовать прочитать их и, возможно, даже запомнить парочку!
Предлагаем начать с «supercalifragilisticexpialidocious». И это возможно!
Не верите? Тогда смотрите!
Sometimes a picture says a thousand words; sometimes a word says a thousand letters. There are a few instances in the English language where a word is not constructed for the sake of communication so much as to break a world record, for spectacle’s sake. In that way, the English language is much like the Olympics; here are ten words that really go the distance.
Note: the following are words in the non-strictest sense, being that some are technical terms, some have been coined, while others actually appear in the dictionary. Depending on which school of thought you subscribe to, lists may very on the basis of “what constitutes a word” (and some may argue simply that letters constitute a word).
Additional note: tying for the #7 spot is the word “hippopotomonstrosesquipedalian.” It also contains 30 letters. Let its omission be justified by saying this list, in and of itself, is hippopotomonstrosesquipedalian (i.e. “that which pertains to extremely long words”).
10. Honorificabilitudinitatibus
This 27-letter word coined by Shakespeare, in his comedy Love’s Labour’s Lost, is a testament to the Bard’s own intralexiconic skills. Meaning “the state of being able to achieve honors,” the word is the longest one in the English language with alternating consonants and vowels (Take a look for yourself….yep.).
9. Antidisestablishmentarianism
Containing 28-letters, antidisestablishmentarianism is the longest proper word, consisting of proper and compatible root and affix attachments. After all the Lego blocks have been snapped together, the word comes to mean “the movement or ideology that opposes disestablishment (i.e. the separation of church and state, as in the movement that took place in 1860’s England).” The word has a dated relevance, or else is the greatest living thing in a world history nerd’s vocabulary.
8. Floccinaucinihilipilification
This 29-letter word, pieced together from Latin stems, means simply “the deeming of something to be trivial.” One letter more than antidisestablishmentarianism, and just as big of a mouthful, it is a valid dictionary entry with a usefulness that is much greater than anything it might be placed beside contextually. Some readers might even be able to maintain a floccinaucinihilipilification for this list.
7. Pseudopseudohypoparathyroidism
This 30-letter word is a technical one for a type of inherited disorder. An individual with such a disorder resembles someone with Pseudohypoparathyroidism Type 1A, but doesn’t possess a deficiency in calcium or PTH levels (which mark the essential differences between Pseudohypoparathyroidism 1A and Hypoparathyroidism).
To put it far more basically, the word is much more fun to say than to have.
6. Supercalifragilisticexpialidocious
This 34-letter word, which was coined by song-writers Richard and Robert Sherman in the musical film Mary Poppins, is completely made-up, the sum of word parts that don’t even follow proper prefix/suffix placement protocol; the “-istic” following “fragil-” is a suffix, which should signify a word’s end. However, it is followed by the prefix “ex-,” where a new, separate word should begin. Nonetheless, it is just another example of a phrase being irretrievably carried off by and imbedded within the culture into which it was born. Just as how words are invented all the time in rap culture, and swallowed up by a constantly-evolving (or devolving) language system.
The word, containing definable roots, means something like “Atoning for educability through delicate beauty.” Miss Poppins, however, would insist the word means “something to say when you have nothing to say.”
5. Pneumonoultramicroscopicsilicovolcanoconiosis
The term refers to a kind of lung disease caused by a finely-powdered silica dust. This word, containing 45 letters, does appear in the dictionary, but was created primarily just for the sake of a long word. An equivalent of what the word is going for, albeit by taking the long way home, is a condition simply called silicosis. Any doctor diagnosing the former is obviously getting paid by the hour.
4. Aequeosalinocalcalinoceraceoaluminosocupreovitriolic
https://www.youtube.com/watch?v=OSppSTBqIvU
This 52-letter word was engendered by Dr. Edward Strother in order to describe spa waters of Bath, England in a single word. The sum of individually-meaningful parts, the word altogether means roughly, “equally salty, calcium-rich, waxy, containing aluminum and copper, and vitriolic.” Of course this word has very few applications elsewhere. Unless maybe you were talking about some kind of 9V-battery-powered robotic chicken wing.
3. Lopadotemachoselachogaleokranioleipsanodrimhypotrimmatosilphioparaomelito-katakechymenokichlepikossyphophattoperisteralektryonoptekephalliokigklopeleiolagoio-siraiobaphetraganopterygon
https://youtu.be/9OGEFqV70Ms?t=152
This word, which shan’t be uttered twice, is a transliteration of a word coined by Greek author Aristophanes in his comic play Assemblywomen. Containing 171 letters, it is the longest word appearing in literature and refers to a fictional dish; the word quite literally is just the smooshing together of the 17 ingredients contained within (including sharks, pigeons, honey, and various unappetizing animal parts). Don’t expect to see this listed on any menu, as there surely wouldn’t be enough room to list anything else.
2. Methionylglutaminylarginyltyrosylglutamylserylleucylphenyl-
alanylalanylglutaminylleucyllysylglutamylarginyllysylglutamyl-
glycylalanylphenylalanylvalylprolylphenylalanylvalylthreonyl-
leucylglycylaspartylprolylglycylisoleucylglutamylglutaminyl-
serylleucyllysylisoleucylaspartylthreonylleucylisoleucylglutamy-
lalanylglycylalanylaspartylalanylleucylglutamylleucylglycylisoleucyl-
prolylphenylalanylserylaspartylprolylleucylalanylaspartylglycylproly-
lthreonylisoleucylglutaminylaspfraginylalanylthreonylleucylarginy-
lalanylphenylalanylalanylalanylglycylvalylthreonylprolylalanyl-
glutaminylcysteinylphenylalanylglutamylmethionylleucylalany-
lleucylisoleucylarginylglutaminyllysylhistidylprolylthreonylisoleucyl-
prolylisoleucylglycylleucylleucylmethionyltyrosylalanylasparaginy-
lleucylvalylphenylalanylasparaginyllysylglycylisoleucylaspartyl-glutamylphenylalanyltyrosylalanylglutaminylcysteinylglutamylly-sylvalylglycylvalylaspartylserylvalylleucylvalylalanylaspartylvalyl-prolylvalylglutaminylglutamylserylalanylprolylphenylalanylarginyl-glutaminylalanylalanylleucylarginylhistidylasparaginylvalylalanyl-prolylisoleucylphenylalanylisoleucylcysteinylprolylprolylaspartylalanyl-aspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanyl-seryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylseryl-arginylalanylglycylvalylthreonylglycylalanylglutamylasparaginyl-arginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalyl-alanyllysylleucyllysylglutamyltyrosylasparaginylalanylalanylprolyl-prolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanyl-prolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanyl-glycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisoleucyl-valyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginy-lisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanyl-leucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanyl-alanylthreonylarginylserine
What you just stared at is the 1913-letter chemical name for tryptophan synthetase, a protein (an enzyme, to be exact) with 267 amino acids. Of course, it’s completely impractical to actually utter this prankster’s approach to making huge words (the largest one in print), and just looking at it for too long might even lead you to believe there are words and phrases hidden in there like a word search (if you look closely the word “party” shows up a few times, as does something resembling “asparagus”). Of course, when you cut-and-paste such a word (rather than risk missing even a single letter, for accuracy’s sake), you risk looking like an ass by not thoroughly combing through that contrived brick-o’-letters.
1. [Titin’s Chemical Name]
This 189,819-letter word shall not be printed in its entirety, partially because it is literally too big to print (without filling the space of a short novella that is), and would be a waste of time and hard drive space. Along the same lines as the last example, it is a derivation of the chemical components that comprise the protein; abridged, the word is “Methionylthreonylthreonyl…isoleucine,” really not worth seeing sprawled-out if for a single-purpose novelty (the only real purpose a chemical name could possibly serve).
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A man has been recorded spending more than three hours to pronounce what is supposedly the longest word in the English language.
‘Methionylthreonylthreonylglutaminylarginyl…isoleucine’ is the chemical name of ‘titin’ (also known as ‘connectin’) — the largest known protein. It has 189,819 letters.
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However, in the past it has been disputed whether or not it is a real word — proteins are named after the chemicals involved in making them.
The longest word in the Oxford English Dictionary is ‘pneumonoultramicroscopicsilicovolcanoconiosis’ at 45 letters long.
‘Supercalifragilisticexpialidocious’, made famous by Mary Poppins, is 34 letters long.
> ‘Amazeballs’, ‘mummy porn’, ‘frenemy’ make Collins online dictionary
> OMG, LOL enter Oxford English Dictionary
Below is the full 189,819-lettered word for ‘titin’:
Methionylthreonylthreonylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylglutaminylleuc yllysylglutamylarginyllysylglutamylglycylalanylphenylalanylvalylprolylphenylalanylvalylthreonylleucylgl ycylaspartylprolylglycylisoleucylglutamylglutaminylserylleucyllysylisoleucylaspartylthreonylleucylisoleu cylglutamylalanylglycylalanylaspartylalanylleucylglutamylleucylglycylisoleucylprolylphenylalanylseryla spartylprolylleucylalanylaspartylglycylprolylthreonylisoleucylglutaminylasparaginylalanylthreonylleucyl arginylalanylphenylalanylalanylalanylglycylvalylthreonylprolylalanylglutaminylcysteinylphenylalanylglu tamylmethionylleucylalanylleucylisoleucylarginylglutaminyllysylhistidylprolylthreonylisoleucylprolylisol eucylglycylleucylleucylmethionyltyrosylalanylasparaginylleucylvalylphenylalanylasparaginyllysylglycyli soleucylaspartylglutamylphenylalanyltyrosylalanylglutaminylcysteinylglutamyllysylvalylglycylvalylaspa rtylserylvalylleucylvalylalanylaspartylvalylprolylvalylglutaminylglutamylserylalanylprolylphenylalanylarg inylglutaminylalanylalanylleucylarginylhistidylasparaginylvalylalanylprolylisoleucylphenylalanylisoleuc ylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalany lseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylgly cylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanylly sylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycy lisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylala nylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhist idylasparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylal anylvalylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreo nylserylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartyl prolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylgl utaminylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylval ylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminy lserylthreonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylaspar aginylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalany laspartylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminy lglutaminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylv alylaspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginyl leucylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminyl asparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylala nylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcyst einylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylserylt yrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylala nylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleu cyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisole ucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylgl ycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidyla sparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylv alylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylser ylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolyli soleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutami nylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglut aminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylseryl threonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparagin ylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylasp artylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglut aminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalyl aspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleu cylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasp araginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanyl prolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcystein ylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyr osylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalany lglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyll ysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucy lserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycyl alanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylaspa raginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalyl glutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylp rolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisol eucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminyl phenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutam inylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthr eonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylal anylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylasparty lthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutami nylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylasp artylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylv alylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylaspara ginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprol ylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylpr olylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosyl 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lylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucyl serylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucy lglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylproly lmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylgluta minylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamyll eucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenylalanylgl utaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminyl phenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylargi nylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleucy 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asparaginylglutaminylasparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryp tophylthreonylserylalanylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylala nylalanylisoleucylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutamin ylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycyl valylthreonylglycylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleu cylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylph enylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalany lglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutam ylglutaminylhistidylasparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllys 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ylarginylarginylvalylaspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisol eucylasparaginylleucylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylaspar aginylglutaminylasparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophy lthreonylserylalanylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylal anylmethionylthreonylthreonylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylglutaminyll eucyllysylglutamylarginyllysylglutamylglycylalanylphenylalanylvalylprolylphenylalanylvalylthreonylleuc ylglycylaspartylprolylglycylisoleucylglutamylglutaminylserylleucyllysylisoleucylaspartylthreonylleucylis oleucylglutamylalanylglycylalanylaspartylalanylleucylglutamylleucylglycylisoleucylprolylphenylalanylse rylaspartylprolylleucylalanylaspartylglycylprolylthreonylisoleucylglutaminylasparaginylalanylthreonylle ucylarginylalanylphenylalanylalanylalanylglycylvalylthreonylprolylalanylglutaminylcysteinylphenylalany lglutamylmethionylleucylalanylleucylisoleucylarginylglutaminyllysylhistidylprolylthreonylisoleucylprolyli soleucylglycylleucylleucylmethionyltyrosylalanylasparaginylleucylvalylphenylalanylasparaginyllysylgly cylisoleucylaspartylglutamylphenylalanyltyrosylalanylglutaminylcysteinylglutamyllysylvalylglycylvalyla spartylserylvalylleucylvalylalanylaspartylvalylprolylvalylglutaminylglutamylserylalanylprolylphenylalany larginylglutaminylalanylalanylleucylarginylhistidylasparaginylvalylalanylprolylisoleucylphenylalanylisol eucylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylal anylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreony lglycylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalan yllysylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylgl ycylisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalany lalanylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminyl histidylasparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphen ylalanylvalylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylth reonylserylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspa rtylprolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparagin ylglutaminylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminy lvalylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylgluta minylserylthreonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosyl asparaginylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenyl alanylaspartylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglut aminylglutaminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginyla rginylvalylaspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylaspa raginylleucylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglut aminylasparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonyls erylalanylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleu cylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalan ylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylgl ycylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyll ysylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglyc ylisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylal anylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhi stidylasparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenyl alanylvalylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthre onylserylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspart ylprolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginyl glutaminylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylv alylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutami nylserylthreonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylas paraginylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylal anylaspartylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylgluta minylglutaminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylar ginylvalylaspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylaspar aginylleucylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylgluta minylasparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylse rylalanylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleuc ylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalany lseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylgly cylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanylly sylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycy lisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylala nylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhist idylasparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylal anylvalylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreo nylserylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartyl prolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylgl utaminylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylval ylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminy lserylthreonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylaspar aginylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalany laspartylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminy lglutaminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylv alylaspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginyl leucylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminyl asparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylala nylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcyst einylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylserylt yrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylala nylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleu cyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisole ucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylgl ycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidyla sparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylv alylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylser ylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolyli soleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutami nylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglut aminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylseryl threonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparagin ylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylasp artylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglut aminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalyl aspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleu cylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasp araginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanyl prolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylmethionylthreon ylthreonylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylglutaminylleucyllysylglutamylar ginyllysylglutamylglycylalanylphenylalanylvalylprolylphenylalanylvalylthreonylleucylglycylaspartylprolyl glycylisoleucylglutamylglutaminylserylleucyllysylisoleucylaspartylthreonylleucylisoleucylglutamylalanyl glycylalanylaspartylalanylleucylglutamylleucylglycylisoleucylprolylphenylalanylserylaspartylprolylleucy lalanylaspartylglycylprolylthreonylisoleucylglutaminylasparaginylalanylthreonylleucylarginylalanylphen ylalanylalanylalanylglycylvalylthreonylprolylalanylglutaminylcysteinylphenylalanylglutamylmethionylleu cylalanylleucylisoleucylarginylglutaminyllysylhistidylprolylthreonylisoleucylprolylisoleucylglycylleucylle ucylmethionyltyrosylalanylasparaginylleucylvalylphenylalanylasparaginyllysylglycylisoleucylaspartylgl utamylphenylalanyltyrosylalanylglutaminylcysteinylglutamyllysylvalylglycylvalylaspartylserylvalylleucyl valylalanylaspartylvalylprolylvalylglutaminylglutamylserylalanylprolylphenylalanylarginylglutaminylalan ylalanylleucylarginylhistidylasparaginylvalylalanylprolylisoleucylphenylalanylisoleucylcysteinylprolylpr olylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycyl arginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla spartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalyla sparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylt hreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylala nyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylp rolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycy larginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla spartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalyla sparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylt hreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylala nyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylp rolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycy larginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla spartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalyla sparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylt hreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylala nyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylp rolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycy larginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla spartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalyla sparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylt hreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylala nyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylmethionylthreonylthreon ylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylglutaminylleucyllysylglutamylarginyllys ylglutamylglycylalanylphenylalanylvalylprolylphenylalanylvalylthreonylleucylglycylaspartylprolylglycylis oleucylglutamylglutaminylserylleucyllysylisoleucylaspartylthreonylleucylisoleucylglutamylalanylglycylal 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ylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylalanyltitinmethionylglutaminylarginy ltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylprolylaspartylalanylaspartylaspart ylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonyltyros ylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginylarginylalanylalanylle ucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanylalan ylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalylly sylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisole ucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylprolylglutamyllys ylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanylalan 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anylmethionylthreonylthreonylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylglutaminyll eucyllysylglutamylarginyllysylglutamylglycylalanylphenylalanylvalylprolylphenylalanylvalylthreonylleuc ylglycylaspartylprolylglycylisoleucylglutamylglutaminylserylleucyllysylisoleucylaspartylthreonylleucylis oleucylglutamylalanylglycylalanylaspartylalanylleucylglutamylleucylglycylisoleucylprolylphenylalanylse rylaspartylprolylleucylalanylaspartylglycylprolylthreonylisoleucylglutaminylasparaginylalanylthreonylle ucylarginylalanylphenylalanylalanylalanylglycylvalylthreonylprolylalanylglutaminylcysteinylphenylalany lglutamylmethionylleucylalanylleucylisoleucylarginylglutaminyllysylhistidylprolylthreonylisoleucylprolyli soleucylglycylleucylleucylmethionyltyrosylalanylasparaginylleucylvalylphenylalanylasparaginyllysylgly cylisoleucylaspartylglutamylphenylalanyltyrosylalanylglutaminylcysteinylglutamyllysylvalylglycylvalyla 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ylalanylvalylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylth reonylserylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspa rtylprolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparagin ylglutaminylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminy lvalylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylgluta minylserylthreonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosyl asparaginylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenyl alanylaspartylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglut aminylglutaminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginyla rginylvalylaspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylaspa raginylleucylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglut aminylasparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonyls erylalanylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleu cylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalan ylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylgl ycylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyll ysylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglyc ylisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylal anylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhi stidylasparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenyl alanylvalylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthre onylserylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspart ylprolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginyl glutaminylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylv alylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutami nylserylthreonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylas paraginylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylal anylaspartylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylgluta minylglutaminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylar ginylvalylaspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylaspar aginylleucylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylgluta minylasparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylse rylalanylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleuc ylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalany lseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylgly cylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanylly sylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycy lisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylala nylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhist idylasparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylal anylvalylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreo nylserylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartyl prolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylgl utaminylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylval ylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminy lserylthreonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylaspar aginylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalany laspartylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminy lglutaminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylv alylaspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginyl leucylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminyl asparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylala nylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcyst einylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylserylt yrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylala nylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleu cyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisole ucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylgl ycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidyla sparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylv alylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylser ylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolyli soleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutami nylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglut aminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylseryl threonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparagin ylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylasp artylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglut aminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalyl aspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleu cylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasp araginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanyl prolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylmethionylthreon ylthreonylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylglutaminylleucyllysylglutamylar ginyllysylglutamylglycylalanylphenylalanylvalylprolylphenylalanylvalylthreonylleucylglycylaspartylprolyl glycylisoleucylglutamylglutaminylserylleucyllysylisoleucylaspartylthreonylleucylisoleucylglutamylalanyl glycylalanylaspartylalanylleucylglutamylleucylglycylisoleucylprolylphenylalanylserylaspartylprolylleucy lalanylaspartylglycylprolylthreonylisoleucylglutaminylasparaginylalanylthreonylleucylarginylalanylphen ylalanylalanylalanylglycylvalylthreonylprolylalanylglutaminylcysteinylphenylalanylglutamylmethionylleu cylalanylleucylisoleucylarginylglutaminyllysylhistidylprolylthreonylisoleucylprolylisoleucylglycylleucylle ucylmethionyltyrosylalanylasparaginylleucylvalylphenylalanylasparaginyllysylglycylisoleucylaspartylgl utamylphenylalanyltyrosylalanylglutaminylcysteinylglutamyllysylvalylglycylvalylaspartylserylvalylleucyl valylalanylaspartylvalylprolylvalylglutaminylglutamylserylalanylprolylphenylalanylarginylglutaminylalan ylalanylleucylarginylhistidylasparaginylvalylalanylprolylisoleucylphenylalanylisoleucylcysteinylprolylpr olylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycyl arginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla spartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalyla sparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylt hreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylala nyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylp rolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycy larginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla spartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalyla sparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylt hreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylala nyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylp rolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycy larginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla spartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalyla sparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylt hreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylala nyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylp rolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycy larginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla spartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalyla sparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylt hreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylala nyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylmethionylthreonylthreon ylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylglutaminylleucyllysylglutamylarginyllys ylglutamylglycylalanylphenylalanylvalylprolylphenylalanylvalylthreonylleucylglycylaspartylprolylglycylis oleucylglutamylglutaminylserylleucyllysylisoleucylaspartylthreonylleucylisoleucylglutamylalanylglycylal anylaspartylalanylleucylglutamylleucylglycylisoleucylprolylphenylalanylserylaspartylprolylleucylalanyla spartylglycylprolylthreonylisoleucylglutaminylasparaginylalanylthreonylleucylarginylalanylphenylalanyl alanylalanylglycylvalylthreonylprolylalanylglutaminylcysteinylphenylalanylglutamylmethionylleucylalan ylleucylisoleucylarginylglutaminyllysylhistidylprolylthreonylisoleucylprolylisoleucylglycylleucylleucylmet hionyltyrosylalanylasparaginylleucylvalylphenylalanylasparaginyllysylglycylisoleucylaspartylglutamylp henylalanyltyrosylalanylglutaminylcysteinylglutamyllysylvalylglycylvalylaspartylserylvalylleucylvalylala nylaspartylvalylprolylvalylglutaminylglutamylserylalanylprolylphenylalanylarginylglutaminylalanylalanyl leucylarginylhistidylasparaginylvalylalanylprolylisoleucylphenylalanylisoleucylcysteinylprolylprolylaspa rtylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylgl ycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparagi nylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyros ylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolyl aspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylse rylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylgl utamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylm ethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutamin ylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleuc ylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenylalanylgluta minylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphe nylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylarginylp henylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleucylasp artylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreonylarginylas paraginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylserylprolylthre onylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartylaspartylalany lthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalylasparaginylgl utamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylthreonylphe nylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylalanyltitinmethi onylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylprolylaspartyl alanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycy ltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginyl arginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosyla 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utaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylprolylaspartylalanyl aspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyros ylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginylarginy lalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylaspara ginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylg lutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycyls erylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylpr olylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyll ysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenyl alanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleucyl asparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenylalanylglutaminylth reonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphenylalan ylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylarginylphenylal anylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleucylaspartylpr olylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreonylarginylasparagi nylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylserylprolylthreonylthr eonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartylaspartylalanylthreon ylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalylasparaginylglutamyll 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ucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanylalan ylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalylly sylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisole ucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylprolylglutamyllys ylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanylalan ylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenylalanylvalylphe nylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleucylasparaginylval ylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenylalanylglutaminylthreonylglutamin ylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphenylalanylserylglutamin 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aspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosyl leucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginylarginylalanylalanylleuc ylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanylalanyl prolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalyllys ylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisoleu cylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylprolylglutamyllysyl methionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanylalanylt hreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenylalanylvalylpheny lalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleucylasparaginylvalylc 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onylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenylalanylvalylphenylala nylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleucylasparaginylvalylcyst einylthreonylserylserylleucylglycylasparaginylglutaminylphenylalanylglutaminylthreonylglutaminylgluta minylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphenylalanylserylglutaminylvalyl tryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylarginylphenylalanylprolylglycylaspar tylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleucylaspartylprolylleucylisoleucylthre onylalanylleucylleucylglycylthreonylphenylalanylaspartylthreonylarginylasparaginylarginylisoleucylisol eucylglutamylvalylglutamylasparaginylglutaminylglutaminylserylprolylthreonylthreonylalanylglutamylth reonylleucylaspartylalanylthreonylarginylarginylvalylaspartylaspartylalanylthreonylvalylalanylisoleucyl 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anylglutaminylcysteinylphenylalanylglutamylmethionylleucylalanylleucylisoleucylarginylglutaminyllysyl histidylprolylthreonylisoleucylprolylisoleucylglycylleucylleucylmethionyltyrosylalanylasparaginylleucylv alylphenylalanylasparaginyllysylglycylisoleucylaspartylglutamylphenylalanyltyrosylalanylglutaminylcys teinylglutamyllysylvalylglycylvalylaspartylserylvalylleucylvalylalanylaspartylvalylprolylvalylglutaminylgl utamylserylalanylprolylphenylalanylarginylglutaminylalanylalanylleucylarginylhistidylasparaginylvalylal anylprolylisoleucylphenylalanylisoleucylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleuc ylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucyl serylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginylarginylalanylalanylleucylprolylleuc ylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolyll 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cylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalan ylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylgl ycylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyll ysylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglyc ylisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylal anylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhi stidylasparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenyl alanylvalylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthre onylserylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspart ylprolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginyl glutaminylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylv alylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutami nylserylthreonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylas paraginylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylal anylaspartylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylgluta minylglutaminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylar ginylvalylaspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylaspar aginylleucylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylgluta minylasparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylse rylalanylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleuc ylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalany lseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylgly cylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanylly sylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycy lisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylala nylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhist idylasparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylal anylvalylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreo nylserylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartyl prolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylgl utaminylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylval ylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminy lserylthreonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylaspar aginylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalany laspartylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminy lglutaminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylv alylaspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginyl leucylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminyl asparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylala nylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcyst einylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylserylt yrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylala nylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleu cyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisole ucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylgl ycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidyla sparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylv alylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylser ylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolyli soleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutami nylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglut aminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylseryl threonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparagin ylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylasp artylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglut aminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalyl aspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleu cylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasp araginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanyl prolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylmethionylthreon ylthreonylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylglutaminylleucyllysylglutamylar ginyllysylglutamylglycylalanylphenylalanylvalylprolylphenylalanylvalylthreonylleucylglycylaspartylprolyl glycylisoleucylglutamylglutaminylserylleucyllysylisoleucylaspartylthreonylleucylisoleucylglutamylalanyl glycylalanylaspartylalanylleucylglutamylleucylglycylisoleucylprolylphenylalanylserylaspartylprolylleucy lalanylaspartylglycylprolylthreonylisoleucylglutaminylasparaginylalanylthreonylleucylarginylalanylphen ylalanylalanylalanylglycylvalylthreonylprolylalanylglutaminylcysteinylphenylalanylglutamylmethionylleu cylalanylleucylisoleucylarginylglutaminyllysylhistidylprolylthreonylisoleucylprolylisoleucylglycylleucylle ucylmethionyltyrosylalanylasparaginylleucylvalylphenylalanylasparaginyllysylglycylisoleucylaspartylgl utamylphenylalanyltyrosylalanylglutaminylcysteinylglutamyllysylvalylglycylvalylaspartylserylvalylleucyl valylalanylaspartylvalylprolylvalylglutaminylglutamylserylalanylprolylphenylalanylarginylglutaminylalan ylalanylleucylarginylhistidylasparaginylvalylalanylprolylisoleucylphenylalanylisoleucylcysteinylprolylpr olylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycyl arginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla spartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalyla sparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylt hreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylala nyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylp rolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycy larginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla spartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalyla sparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylt hreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylala nyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylp rolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycy larginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla spartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalyla sparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylt hreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylala nyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylp rolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycy larginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala 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lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla spartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalyla sparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylt hreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylala nyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylmethionylthreonylthreon ylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylglutaminylleucyllysylglutamylarginyllys ylglutamylglycylalanylphenylalanylvalylprolylphenylalanylvalylthreonylleucylglycylaspartylprolylglycylis 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rtylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylgl ycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparagi nylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyros ylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolyl aspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylse rylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylgl utamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylm ethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutamin ylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleuc 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It should come as no surprise that we are word lovers. In fact, we are big word lovers in that we love really big words. To express our love, we looked around for some of the biggest, most ludicrously long words in the English language. In addition to pure length, we also tried to find:
- the longest word without vowels
- the longest one-syllable word
- and other uniquely long words.
Figuring out which word is the longest of them all isn’t as simple as just counting letters, though. Should you count scientific words? Should obscure, rarely used words be included, or should we give the honor to a word people actually use? In the interest of fairness, our list includes scientific words, obscure words, and all of the absurdly long words stuck in between.
methionylthreonylthreonylglutaminylarginyl…
At over 180,000 letters long, the chemical name of the protein titin is often said to technically be the longest English word. If spoken out loud, this word takes over three hours to say! Its absurd length is due to the fact that proteins get their scientific names by combining the names of all of their joined amino acids together, and titin has quite a lot of them. For obvious reasons, titin’s official name has never actually appeared in a dictionary or scientific text. Because it is a scientific term, many would disqualify the Big M from actually taking the crown as English’s longest word.
pneumonoultramicroscopicsilicovolcanoconiosis
Pneumonoultramicroscopicsilicovolcanoconiosis, coming in at 45 letters long, is typically the biggest word you will find that actually appears in an English dictionary. According to many sources, it was coined around 1935 by Everett Smith, who at the time was the president of the National Puzzlers’ League. The word, which was basically engineered for its length, refers to a lung disease caused by inhaling silica dust.
sesquipedalianism
Let’s look at a word related to the business of “longest words.” Sesquipedalianism is the tendency to use long words. Do you have sesquipedalian tendencies? (We do.)
The word is traced to the ancient Roman poet Horace, who in a treatise on the art of poetry wrote that in certain circumstances, poets must avoid sesquipedalia verba, a Latin phrase meaning “words [verba] a foot and a half long [sesquipedalia].” Horace clearly had a sense of humor.
pseudopseudohypoparathyroidism
We include pseudopseudohypoparathyroidism, another medical term, because this is one of the longest words to appear in major dictionaries that wasn’t created with length in mind. Pseudo- is a combining form meaning “false, pretended, unreal.”
You might notice the appearance of pseudo- twice. That’s because this disorder simulates the symptoms of pseudohypoparathyroidism, in which the body doesn’t respond to the parathyroid hormone. So, there are two levels of “faking it” going on here.
English isn’t the only language with lengthy elements of its lexicon. Get to know some of the world’s longest words!
antidisestablishmentarianism
Often, people will bust out antidisestablishmentarianism as the longest word they know and are actually able to say. This word has rarely been used and is only mentioned due to its ridiculous length. This word refers to opposition to withdrawing support from the Anglican Church as the state church of 19th-century England.
supercalifragilisticexpialidocious
Even though the sound if it is something quite atrocious, we do really like the word supercalifragilisticexpialidocious. This nonsensical word with no real meaning was popularized by the 1964 Disney film Mary Poppins and is often used by children as an example of a humorously long word. Songwriters Richard and Robert Sherman take credit for this exact spelling of the word, but the word itself existed even before Mary Poppins made it popular.
floccinaucinihilipilification
Here’s one that is also very meta: floccinaucinihilipilification is a rarely used word that means “the estimation of something as valueless.” It is usually used in reference to itself! Dating back to the 1700s, the word contains four Latin roots that all mean “of little value” or “trifling”: floccī, naucī, nihilī, and pilī.
honorificabilitudinitatibus
The word honorificabilitudinitatibus, which is said to mean “capable of receiving honor,” has two major honors to its name. Firstly, it is the longest word to ever appear in the works of William Shakespeare. Billy the Bard only ever used it once, in his play Love’s Labour’s Lost (1590s). Secondly, honorificabilitudinitatibus is the longest English word wherein the consonants and vowels alternate back and forth. Check it again if you didn’t notice!
uncharacteristically
Uncharacteristically for most of the really long words you’ve seen so far, the word uncharacteristically is often said to be the longest word that the average English speaker will commonly see or actually use in everyday life. As you may already know, uncharacteristically is an adverb that describes something as not being typical or acting in a characteristic manner.
incomprehensibilities
At 21 characters, another one of the longest words you might actually use yourself is incomprehensibilities. We define incomprehensible as “impossible to comprehend or understand,” so incomprehensibilities are “things you can’t comprehend.” ¿Comprendes?
uncopyrightables
If you look closely at the spelling, you’ll notice a peculiar thing about this word with 16 letters. It does not repeat any letter; each character is used only once. This word is sometimes called an isogram among lovers of words and word games.
One of the longest isograms is subdermatoglyphic, at 17 characters. But, since subdermatoglyphic (dermatoglyphics studies the patterns of skin markings on the hands and feet) is a bit scientific and certainly not one that is used often, we’re spotlighting uncopyrightables instead because it’s one we can all remember. It means, of course, “items that are unable to be copyrighted.”
rhythms
The word rhythms may not look like much at only seven letters long, but it is said to be the longest English word without one of the five main vowels in it. The letter Y, that wishy-washy “sometimes vowel,” is filling in while A, E, I, O, and U are taking a break. As we all know, the word rhythms means “movements or procedures with uniform or patterned recurrence of a beat, accent, or the like.”
Are there really words without vowels? Depending on the definition, the answer might elicit a hmm.
strengths
Strengths is another smaller word with a big achievement under its belt. It is the longest English word with only a single vowel in it. Considering it is only eight letters long, that really shows you how much we value our vowels. The plural strengths is most often used to mean “positive or valuable attributes or qualities.”
squirrelled
We go nuts for this word. While the word squirreled usually only has one L in American English, some dictionaries accept this British English version as an alternate spelling. Some Americans pronounce the word squirrelled as a one-syllable word (rhyming with “curled”). This makes squirrelled the longest one-syllable word in the English language at 11 letters. If all of that sounds too squirrely for you, the one-syllable verb broughammed from the noun brougham, a type of carriage, is also 11 letters long.
As an aside, some may tell you that the word schtroumpfed is actually the longest “English” word with only a single syllable. This word has been used in some translations of The Smurfs (which is Les Schtroumpfs in French) in place of the more commonly used nonsensical verb smurfed. In our opinion, claiming schtroumpfed is the longest one-syllable word is just a bunch of smurf.
Aegilops
Aegilops sounds like the name of a mythical monster, but it is actually the name of a genus of wild grasses commonly known as goatgrass. In addition to that, Aegilops is also commonly said to be the longest English word that has all of the letters in alphabetical order.
Indulge your inner sesquipedalian and take the quiz!
Think you’re ready to ace our longest English words quiz? Or is it a long shot? It won’t be long before you find out. Remember that a little practice goes a long way!
Have you ever wondered what the longest English word in the world is? Or how many letters does such a word contain? The identity of the longest word in English language could differ by defining a word differently. It could be a notable long word like a place name/personal name or creations of long words such as a coinage/technical term.
What is the Longest Word in the English Language?
What is the longest word in the world? Long words consisting of hundreds of thousands of letters only exist in alphabetic languages like English. For character-based languages like Chinese, a word is made up of one or a few characters, making their length notably limited.
Arbitrary
The length of the English word is most commonly based on orthography and the total number of its written letters. There’s no definite answer to what the world’s longest word is. To give a particularly vivid example, the word “great-great-great-…great-great-grandparent” contains an arbitrary amount of letters, depending on how many “-great”s are attached to the root word “parent.” In such case, no other English word can beat this word in terms of the counting numbers of the written letters because you can add as many “-great”s as you like to extend the length.
Chemical
When it comes to the biggest word in the world by magnitude, the word for the chemical composition of titin (Methionylthreonylthreonylglutaminylalanyl…isoleucine) takes the crown with 189,819 letters. Yes, this long word looks like the text produced by someone who had accidentally sat on the keyboard for a considerable amount of time. If you have three hours to spare and the insane patience, you can try to say the entire word.
Its incredible length and the controversy over whether it should be considered a word have hindered this technical term for the largest known protein from entering the dictionary.
The second longest word in English is Methionylglutaminylarginyltyrosylglutamyl…serine with 1,909 letters. Compared to the longest word in English, this chemical term of E coli TrpA (P0A877) is luckier since it is the longest published word in English, though not in a dictionary.
Ten Longest English Words in the Dictionary
The length of a word is measured by the number of its written letters. In the following passage, we will list the ten longest words in the English dictionaries according to their length. Depending on their usage and popularity, these words are listed in one or more of the major dictionaries.
Forty-five Letters
Pneumonoultramicroscopicsilicovolcanoconiosis, the synonym for the disease silicosis, is the longest word in the most trusted English dictionaries like the Webster’s, Oxford, Chambers, and Random House with 45 letters. This medical term was coined in 1935 and describes the occupational lung disease caused by breathing in crystalline silica dust. Interestingly, this word was purposely designed to invent the then-new longest word. Consequently, the Oxford English Dictionary describes it as a factitious word.
Thirty-seven Letters
The second longest word in the English dictionary is the 37-letters-long word hepaticocholangiocholecystenterostomies, a surgical term in Gould’s Medical Dictionary that describes it as the creation of a link between a hepatic and the gall bladder and between the gall bladder and the intestine.
Thirty-four Letters
The third longest word published in the English dictionary is supercalifragilisticexpialidocious with 34 letters. It is a song and single by Julie Andrews and Dick Van Dyke from the Disney musical film Mary Poppins in 1964.
Thirty Letters
The adjective hippopotomonstrosesquipedalian is 30 letters long, ranking fourth on our list of the longest English words in the dictionary. Mrs. Byrne’s Dictionary of Unusual, Obscure and Preposterous Words defines it as of/pertaining to an extremely long word.
Twenty-nine Letters
The word floccinaucinihilipilification comes fifth with 29 letters. Dating back to 1741, it is the longest word in the 1st edition of the Oxford English Dictionary and the 1992 Guinness Book of World Records refers to it as the longest real word. This uncountable noun is the act/habit of regarding something as worthless/unimportant. Interestingly, the common English letter E doesn’t occur in this word, while the letter I appears nine times in total.
Twenty-eight Letters
The sixth longest word in the English dictionary is antidisestablishmentarianism, with 28 letters. It refers to a political philosophy that opposes the withdrawal of state recognition or support from its national Church. This word is considered one the most popular longest words in English in the past decades.
Twenty-seven Letters
Honorificabilitudinitatibus, the synonym for honorableness, comes seventh on the long English words list, with two other 27-letters-long words being electroencephalographically and antitransubstantiationalist. It means the state of being able to achieve honors. Honorificabilitudinitatibus first appeared as an English word in 1599 and entered Bailey’s Dictionary in 1721 as the longest word in English back then. Additionally, it is the longest English word Shakespeare ever used in his works.
Electroencephalographically is tied for the seventh longest word in English dictionary, meaning by means of electroencephalography, an apparatus that detects electrical potentials on the scalp and records brain waves. It is the longest unhyphenated word listed in the 10th edition of the Merriam-Webster’s Collegiate Dictionary together with the chemical term ethylenediaminetetraacetate, a salt of Ethylene Diamine Tetraacetic Acid.
Another 27-letter word is antitransubstantiationalist, a religious term that describes someone who disagrees that the consecrated bread and wine can actually change into the body and blood of Christ.
Translating the Longest Word in English
The longest word in the English language can come in many forms. It could be Taumatawhakatangihangakoauauotamateaturipukakapikimaungahoronukupokaiwhenuakitanatahu, the 85-letters-long place name for a hill near the town of Porangahau in New Zealand. Or it could be Hubert Blaine Wolfeschlegelsteinhausenbergerdorff Sr., the longest personal name ever used that was made up of 747 characters. Whether it’s the medical word pneumonoultramicroscopicsilicovolcanoconiosis or other incredibly long chemical terms, translating such words will require in-country/industry specific linguists who can guarantee the accuracy of its translations based on their linguistic, cultural, and technical knowledge of the subject matter.
Getting the correct pronunciation of the longest word in English is a Herculean task per se. Not to mention translating it right. That’s why you need to work with professionals. If you need such experts to handle the translation of just about any content from and into English or any other language in the matter, Wordspath is the go-to option! Contact us to further discuss your linguistic needs by hitting the button below.
Conclusion
The longest word in English is not always the same one. As time goes by, new long words will be coined and outrank the old ones on the long words list. In the great world where wonders never cease, which long word lives up to the name of the world’s longest word is really up to your preference.