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The identity of the longest word in English depends on the definition of a word and of length.

Words may be derived naturally from the language’s roots or formed by coinage and construction. Additionally, comparisons are complicated because place names may be considered words, technical terms may be arbitrarily long, and the addition of suffixes and prefixes may extend the length of words to create grammatically correct but unused or novel words.

The length of a word may also be understood in multiple ways. Most commonly, length is based on orthography (conventional spelling rules) and counting the number of written letters. Alternate, but less common, approaches include phonology (the spoken language) and the number of phonemes (sounds).

Word Letters Meaning Claim Dispute
methionylthreonylthreonylglutaminylalanyl…isoleucine 189,819 The chemical composition of titin, the largest known protein Longest known word overall by magnitudes. Attempts to say the entire word have taken two[1] to three and a half hours.[2] Technical; not in dictionary; whether this should actually be considered a word is disputed
methionylglutaminylarginyltyrosylglutamyl…serine 1,909 The chemical name of E. coli TrpA (P0A877) Longest published word[3] Technical
lopadotemachoselachogaleokranioleipsano…pterygon 183 A fictional dish of food Longest word coined by a major author,[4] the longest word ever to appear in literature[5] Contrived nonce word; not in dictionary; Ancient Greek transliteration
pneumonoultramicroscopicsilicovolcanoconiosis 45 The disease silicosis Longest word in a major dictionary[6] Contrived coinage to make it the longest word; technical, but only mentioned and never actually used in communication
supercalifragilisticexpialidocious 34 Unclear in source work, has been cited as a nonsense word Made popular in the Mary Poppins film and musical[7] Contrived coinage
pseudopseudohypoparathyroidism 30 A hereditary medical disorder Longest non-contrived word in a major dictionary[8] Technical
antidisestablishmentarianism 28 The political position of opposing disestablishment Longest non-contrived and nontechnical word[9] Not all dictionaries accept it due to lack of usage.[10]
honorificabilitudinitatibus 27 The state of being able to achieve honors Longest word in Shakespeare’s works; longest word in the English language featuring alternating consonants and vowels[11] Latin

Major dictionaries

The longest word in any of the major English language dictionaries is pneumonoultramicroscopicsilicovolcanoconiosis (45 letters), a word that refers to a lung disease contracted from the inhalation of very fine silica particles,[12] specifically from a volcano; medically, it is the same as silicosis. The word was deliberately coined to be the longest word in English, and has since been used[citation needed] in a close approximation of its originally intended meaning, lending at least some degree of validity to its claim.[6]

The Oxford English Dictionary contains pseudopseudohypoparathyroidism (30 letters).

Merriam-Webster’s Collegiate Dictionary does not contain antidisestablishmentarianism (28 letters), as the editors found no widespread, sustained usage of the word in its original meaning. The longest word in that dictionary is electroencephalographically (27 letters).[13]

The longest non-technical word in major dictionaries is flocci­nauci­nihili­pili­fication at 29 letters. Consisting of a series of Latin words meaning «nothing» and defined as «the act of estimating something as worthless»; its usage has been recorded as far back as 1741.[14][15][16]

Ross Eckler has noted that most of the longest English words are not likely to occur in general text, meaning non-technical present-day text seen by casual readers, in which the author did not specifically intend to use an unusually long word. According to Eckler, the longest words likely to be encountered in general text are deinstitutionalization and counterrevolutionaries, with 22 letters each.[17]

A computer study of over a million samples of normal English prose found that the longest word one is likely to encounter on an everyday basis is uncharacteristically, at 20 letters.[18]

The word internationalization is abbreviated «i18n», the embedded number representing the number of letters between the first and the last.[19][20][21]

Creations of long words

Coinages

In his play Assemblywomen (Ecclesiazousae), the ancient Greek comedic playwright Aristophanes created a word of 171 letters (183 in the transliteration below), which describes a dish by stringing together its ingredients:

Henry Carey’s farce Chrononhotonthologos (1743) holds the opening line: «Aldiborontiphoscophornio! Where left you Chrononhotonthologos?»

Thomas Love Peacock put these creations into the mouth of the phrenologist Mr. Cranium in his 1816 book Headlong Hall: osteosarchaematosplanchnochondroneuromuelous (44 characters) and osseocarnisanguineoviscericartilaginonervomedullary (51 characters).

James Joyce made up nine 100-letter words plus one 101-letter word in his novel Finnegans Wake, the most famous of which is Bababadalgharaghtakamminarronnkonnbronntonnerronntuonnthunntrovarrhounawnskawntoohoohoordenenthurnuk. Appearing on the first page, it allegedly represents the symbolic thunderclap associated with the fall of Adam and Eve. As it appears nowhere else except in reference to this passage, it is generally not accepted as a real word. Sylvia Plath made mention of it in her semi-autobiographical novel The Bell Jar, when the protagonist was reading Finnegans Wake.

«Supercalifragilisticexpialidocious», the 34-letter title of a song from the movie Mary Poppins, does appear in several dictionaries, but only as a proper noun defined in reference to the song title. The attributed meaning is «a word that you say when you don’t know what to say.» The idea and invention of the word is credited to songwriters Robert and Richard Sherman.

Agglutinative constructions

The English language permits the legitimate extension of existing words to serve new purposes by the addition of prefixes and suffixes. This is sometimes referred to as agglutinative construction. This process can create arbitrarily long words: for example, the prefixes pseudo (false, spurious) and anti (against, opposed to) can be added as many times as desired. More familiarly, the addition of numerous «great»s to a relative, such as «great-great-great-great-grandparent», can produce words of arbitrary length. In musical notation, an 8192nd note may be called a semihemidemisemihemidemisemihemidemisemiquaver.

Antidisestablishmentarianism is the longest common example of a word formed by agglutinative construction.

Technical terms

A number of scientific naming schemes can be used to generate arbitrarily long words.

The IUPAC nomenclature for organic chemical compounds is open-ended, giving rise to the 189,819-letter chemical name Methionylthreonylthreonyl…isoleucine for the protein also known as titin, which is involved in striated muscle formation. In nature, DNA molecules can be much bigger than protein molecules and therefore potentially be referred to with much longer chemical names. For example, the wheat chromosome 3B contains almost 1 billion base pairs,[22] so the sequence of one of its strands, if written out in full like Adenilyladenilylguanilylcystidylthymidyl…, would be about 8 billion letters long. The longest published word, Acetylseryltyrosylseryliso…serine, referring to the coat protein of a certain strain of tobacco mosaic virus (P03575), is 1,185 letters long, and appeared in the American Chemical Society’s Chemical Abstracts Service in 1964 and 1966.[23] In 1965, the Chemical Abstracts Service overhauled its naming system and started discouraging excessively long names. In 2011, a dictionary broke this record with a 1909-letter word describing the trpA protein (P0A877).[3]

John Horton Conway and Landon Curt Noll developed an open-ended system for naming powers of 10, in which one sexmilliaquingentsexagintillion, coming from the Latin name for 6560, is the name for 103(6560+1) = 1019683. Under the long number scale, it would be 106(6560) = 1039360.

Gammaracanthuskytodermogammarus loricatobaicalensis is sometimes cited as the longest binomial name—it is a kind of amphipod. However, this name, proposed by B. Dybowski, was invalidated by the International Code of Zoological Nomenclature in 1929 after being petitioned by Mary J. Rathbun to take up the case.[24]

Myxococcus llanfairpwllgwyngyllgogerychwyrndrobwllllantysiliogogogochensis is the longest accepted binomial name for an organism. It is a bacterium found in soil collected at Llan­fair­pwll­gwyn­gyll­ (discussed below). Parastratiosphecomyia stratiosphecomyioides is the longest accepted binomial name for any animal, or any organism visible with the naked eye. It is a species of soldier fly.[25] The genus name Parapropalaehoplophorus (a fossil glyptodont, an extinct family of mammals related to armadillos) is two letters longer, but does not contain a similarly long species name.

Aequeosalinocalcalinoceraceoaluminosocupreovitriolic, at 52 letters, describing the spa waters at Bath, England, is attributed to Dr. Edward Strother (1675–1737).[26] The word is composed of the following elements:

  • Aequeo: equal (Latin, aequo[27])
  • Salino: containing salt (Latin, salinus)
  • Calcalino: calcium (Latin, calx)
  • Ceraceo: waxy (Latin, cera)
  • Aluminoso: alumina (Latin)
  • Cupreo: from «copper»
  • Vitriolic: resembling vitriol

Notable long words

Place names

The longest officially recognized place name in an English-speaking country is Taumata­whakatangihanga­koauau­o­tamatea­turi­pukaka­piki­maunga­horo­nuku­pokai­whenua­ki­tana­tahu (85 letters), which is a hill in New Zealand. The name is in the Māori language. A widely recognized version of the name is Taumatawhakatangihangakoauauotamateaturipukakapikimaungahoronukupokaiwhenuakitanatahu (85 letters), which appears on the signpost at the location (see the photo on this page). In Māori, the digraphs ng and wh are each treated as single letters.

In Canada, the longest place name is Dysart, Dudley, Harcourt, Guilford, Harburn, Bruton, Havelock, Eyre and Clyde, a township in Ontario, at 61 letters or 68 non-space characters.[28]

The 58-letter name Llan­fair­pwll­gwyn­gyll­gogery­chwyrn­drob­wlll­lanty­silio­gogo­goch is the name of a town on Anglesey, an island of Wales. In terms of the traditional Welsh alphabet, the name is only 51 letters long, as certain digraphs in Welsh are considered as single letters, for instance ll, ng and ch. It is generally agreed, however, that this invented name, adopted in the mid-19th century, was contrived solely to be the longest name of any town in Britain. The official name of the place is Llanfairpwllgwyngyll, commonly abbreviated to Llanfairpwll or Llanfair PG.

The longest non-contrived place name in the United Kingdom which is a single non-hyphenated word is Cottonshopeburnfoot (19 letters) and the longest which is hyphenated is Sutton-under-Whitestonecliffe (29 characters).

The longest place name in the United States (45 letters) is Char­gogga­gogg­man­chau­ggagogg­chau­buna­gunga­maugg, a lake in Webster, Massachusetts. It means «Fishing Place at the Boundaries – Neutral Meeting Grounds» and is sometimes facetiously translated as «you fish your side of the water, I fish my side of the water, nobody fishes the middle». The lake is also known as Webster Lake.[29] The longest hyphenated names in the U.S. are Winchester-on-the-Severn, a town in Maryland, and Washington-on-the-Brazos, a notable place in Texas history. The longest single-word town names in the U.S. are Kleinfeltersville, Pennsylvania and Mooselookmeguntic, Maine.

The longest official geographical name in Australia is Ma­mungku­kumpu­rang­kunt­junya.[30] It has 26 letters and is a Pitjantjatjara word meaning «where the Devil urinates».[31]

Liechtenstein is the longest country name with single name in English. The second longest country name with single name in English is Turkmenistan. There are longer country names if one includes ones with spaces.

Personal names

Guinness World Records formerly contained a category for longest personal name used.

  • From about 1975 to 1985, the recordholder was Adolph Blaine Charles David Earl Frederick Gerald Hubert Irvin John Kenneth Lloyd Martin Nero Oliver Paul Quincy Randolph Sherman Thomas Uncas Victor William Xerxes Yancy Zeus Wolfe­schlegelstein­hausenberger­dorffvoraltern­waren­gewissenhaft­schaferswessen­schafewaren­wohlgepflege­und­sorgfaltigkeit­beschutzen­von­angreifen­durch­ihrraubgierigfeinde­welche­voraltern­zwolftausend­jahres­vorandieerscheinen­wander­ersteer­dem­enschderraumschiff­gebrauchlicht­als­sein­ursprung­von­kraftgestart­sein­lange­fahrt­hinzwischen­sternartigraum­auf­der­suchenach­diestern­welche­gehabt­bewohnbar­planeten­kreise­drehen­sich­und­wohin­der­neurasse­von­verstandigmen­schlichkeit­konnte­fortplanzen­und­sicher­freuen­anlebens­langlich­freude­und­ruhe­mit­nicht­ein­furcht­vor­angreifen­von­anderer­intelligent­geschopfs­von­hinzwischen­sternartigraum, Senior (746 letters), also known as Wolfe+585, Senior.
  • After 1985 Guinness briefly awarded the record to a newborn girl with a longer name. The category was removed shortly afterward.

Long birth names are often coined in protest of naming laws or for other personal reasons.

  • The naming law in Sweden was challenged by parents Lasse Diding and Elisabeth Hallin, who proposed the given name «Brfxxccxxmnpcccclllmmnprxvclmnckssqlbb11116» for their child (pronounced [ˈǎlːbɪn], 43 characters), which was rejected by a district court in Halmstad, southern Sweden.

Words with certain characteristics of notable length

  • Schmaltzed and strengthed (10 letters) appear to be the longest monosyllabic words recorded in The Oxford English Dictionary, while scraunched and scroonched appear to be the longest monosyllabic words recorded in Webster’s Third New International Dictionary; but squirrelled (11 letters) is the longest if pronounced as one syllable only (as permitted in The Shorter Oxford English Dictionary and Merriam-Webster Online Dictionary at squirrel, and in Longman Pronunciation Dictionary). Schtroumpfed (12 letters) was coined by Umberto Eco, while broughammed (11 letters) was coined by William Harmon after broughamed (10 letters) was coined by George Bernard Shaw.
  • Strengths is the longest word in the English language containing only one vowel letter.[32]
  • Euouae, a medieval musical term, is the longest English word consisting only of vowels, and the word with the most consecutive vowels. However, the «word» itself is simply a mnemonic consisting of the vowels to be sung in the phrase «seculorum Amen» at the end of the lesser doxology. (Although u was often used interchangeably with v, and the variant «Evovae» is occasionally used, the v in these cases would still be a vowel.)
  • The longest words with no repeated letters are dermatoglyphics and uncopyrightable.[33]
  • The longest word whose letters are in alphabetical order is the eight-letter Aegilops, a grass genus. However, this is arguably a proper noun. There are several six-letter English words with their letters in alphabetical order, including abhors, almost, begins, biopsy, chimps and chintz.[34] There are few 7-letter words, such as «billowy» and «beefily». The longest words whose letters are in reverse alphabetical order are sponged, wronged and trollied.
  • The longest words recorded in OED with each vowel only once, and in order, are abstemiously, affectiously, and tragediously (OED). Fracedinously and gravedinously (constructed from adjectives in OED) have thirteen letters; Gadspreciously, constructed from Gadsprecious (in OED), has fourteen letters. Facetiously is among the few other words directly attested in OED with single occurrences of all six vowels (counting y as a vowel).
  • The longest single palindromic word in English is rotavator, another name for a rotary tiller for breaking and aerating soil.

Typed words

  • The longest words typable with only the left hand using conventional hand placement on a QWERTY keyboard are tesseradecades, aftercataracts, dereverberated, dereverberates[35] and the more common but sometimes hyphenated sweaterdresses.[34] Using the right hand alone, the longest word that can be typed is johnny-jump-up, or, excluding hyphens, monimolimnion[36] and phyllophyllin.
  • The longest English word typable using only the top row of letters has 11 letters: rupturewort. The word teetertotter (used in North American English) is longer at 12 letters, although it is usually spelled with a hyphen.
  • The longest using only the middle row is shakalshas (10 letters). Nine-letter words include flagfalls; eight-letter words include galahads and alfalfas.
  • Since the bottom row contains no vowels, no standard words can be formed. [37]
  • The longest words typable by alternating left and right hands are antiskepticism and leucocytozoans respectively.[34]
  • On a Dvorak keyboard, the longest «left-handed» words are epopoeia, jipijapa, peekapoo, and quiaquia.[38] Other such long words are papaya, Kikuyu, opaque, and upkeep.[39] Kikuyu is typed entirely with the index finger, and so the longest one-fingered word on the Dvorak keyboard. There are no vowels on the right-hand side, and so the longest «right-handed» word is crwths.

See also

  • Lipogram
  • List of long species names
  • List of the longest English words with one syllable
  • Longest English sentence
  • Longest word in French
  • Longest word in Romanian
  • Longest word in Spanish
  • Longest word in Turkish
  • Number of words in English
  • Scriptio continua
  • Sesquipedalianism
  • Donau­dampf­schiffahrts­elektrizitäten­haupt­betriebs­werk­bau­unter­beamten­gesellschaft, longest published word in German

References

  1. ^ «Reading The Longest English Word (190,000 Characters)». YouTube. Archived from the original on 2021-11-10. Retrieved 2 August 2020.
  2. ^ «World’s longest word takes 3.5 hours to pronounce». CW39 Houston. 2012-12-08. Retrieved 2020-05-18.
  3. ^ a b Colista Moore (2011). Student’s Dictionary. p. 524. ISBN 978-1-934669-21-1.
  4. ^ see separate article Lopado…pterygon
  5. ^ Donald McFarlan; Norris Dewar McWhirter; David A. Boeh (1989). Guinness book of world records: 1990. Sterling. p. 129. ISBN 978-0-8069-5790-6.
  6. ^ a b Coined around 1935 to be the longest word; press reports on puzzle league members legitimized it somewhat. First appeared in the MWNID supplement, 1939. Today OED and several others list it, but citations are almost always as «longest word». More detail at pneumonoultramicroscopicsilicovolcanoconiosis.
  7. ^ «Merriam Webster: Supercalifragilisticexpialidocious».
  8. ^ «What is the longest English word?». AskOxford. Archived from the original on 2008-10-22. Retrieved 2010-08-22.
  9. ^ «What is the longest English word?». oxforddictionaries.com.[dead link]
  10. ^ «Merriam Webster: «Antidisestablishmentarianism is not in the dictionary.»«.
  11. ^ «Cool, Strange, and Interesting Facts,» fact 99. InnocentEnglish.com. Retrieved 2019-03-13.
  12. ^ «pneumonoultramicroscopicsilicovolcanoconiosis – definition of pneumonoultramicroscopicsilicovolcanoconiosis in English from the Oxford dictionary». oxforddictionaries.com. Archived from the original on 2012-07-19.
  13. ^ «The Longest Word in the Dictionary» (Video). Ask the Editor. Merriam-Webster. Archived from the original on 21 November 2013. Retrieved 14 November 2013.
  14. ^ «Floccinaucinihilipilification» by Michael Quinion World Wide Words Archived 2006-08-21 at the Wayback Machine;
  15. ^ The Guinness Book of Records, in its 1992 and previous editions, declared the longest real word in the English language to be floccinaucinihilipilification. More recent editions of the book have acknowledged pneumonoultramicroscopicsilicovolcanoconiosis. What is the longest English word? — Oxford Dictionaries Online Archived 2006-08-26 at the Wayback Machine
  16. ^ In recent times its usage has been recorded in the proceedings of the United States Senate by Senator Robert Byrd Discussion between Sen. Moynihan and Sen. Byrd «Mr. President, may I say to the distinguished Senator from New York, I used that word on the Senate floor myself 2 or 3 years ago. I cannot remember just when or what the occasion was, but I used it on that occasion to indicate that whatever it was I was discussing it was something like a mere trifle or nothing really being of moment.» Congressional Record June 17, 1991, p. S7887, and at the White House by Bill Clinton’s press secretary Mike McCurry, albeit sarcastically. December 6, 1995, White House Press Briefing in discussing Congressional Budget Office estimates and assumptions: «But if you – as a practical matter of estimating the economy, the difference is not great. There’s a little bit of floccinaucinihilipilification going on here.»
  17. ^ Eckler, R. Making the Alphabet Dance, p 252, 1996.
  18. ^ «Longest Common Words – Modern». Maltron.com. Archived from the original on 27 April 2009. Retrieved 2010-08-22.
  19. ^ «Glossary of W3C Jargon». World Wide Web Consortium. Archived from the original on 2008-10-25. Retrieved 2008-10-13.
  20. ^ «Origin of the Abbreviation I18n». Archived from the original on 2014-06-27.
  21. ^ «Localization vs. Internationalization». World Wide Web Consortium. Archived from the original on 2016-04-03.
  22. ^ Paux et al. (2008) Science, Vol. 322 (5898) 101-104. A Physical Map of the 1-Gigabase Bread Wheat Chromosome 3B Paux, Etienne; Sourdille, Pierre; Salse, Jérôme; Saintenac, Cyrille; Choulet, Frédéric; Leroy, Philippe; Korol, Abraham; Michalak, Monika; Kianian, Shahryar; Spielmeyer, Wolfgang; Lagudah, Evans; Somers, Daryl; Kilian, Andrzej; Alaux, Michael; Vautrin, Sonia; Bergès, Hélène; Eversole, Kellye; Appels, Rudi; Safar, Jan; Simkova, Hana; Dolezel, Jaroslav; Bernard, Michel; Feuillet, Catherine (2008). «A Physical Map of the 1-Gigabase Bread Wheat Chromosome 3B». Science. 322 (5898): 101–104. Bibcode:2008Sci…322..101P. doi:10.1126/science.1161847. PMID 18832645. S2CID 27686615. Archived from the original on 2015-09-03. Retrieved 2012-12-01.
  23. ^ Chemical Abstracts Formula Index, Jan.-June 1964, Page 967F; Chemical Abstracts 7th Coll. Formulas, C23H32-Z, 56-65, 1962–1966, Page 6717F
  24. ^ «Opinion 105. Dybowski’s (1926) Names of Crustacea Suppressed». Opinions Rendered by the International Commission on Zoological Nomenclature: Opinions 105 to 114. Smithsonian Miscellaneous Collections. Vol. 73. 1929. pp. 1–3. hdl:10088/23619. BHL page 8911139.
  25. ^ rjk. «World’s longest name of an animal. Parastratiosphecomyia stratiosphecomyioides Stratiomyid Fly Soldier Fly». thelongestlistofthelongeststuffatthelongestdomainnameatlonglast.com. Archived from the original on 2011-11-17. Retrieved 2011-12-17.
  26. ^ cited in some editions of the Guinness Book of Records as the longest word in English, see Askoxford.com on the longest English word
  27. ^ [1][dead link]
  28. ^ «GeoNames Government of Canada site». Archived from the original on 2009-02-06.
  29. ^ Belluck, Pam (2004-11-20). «What’s the Name of That Lake? It’s Hard to Say». The New York Times.
  30. ^ «Geoscience Australia Gazetteer». Archived from the original on 2007-10-01.
  31. ^ «South Australian State Gazetteer». Archived from the original on 2007-10-01.
  32. ^ «Guinness Records».
  33. ^ «Longest Word Without Repeating Letters». December 2014.
  34. ^ a b c «Typewriter Words». Questrel.com. Archived from the original on 2010-09-27. Retrieved 2010-08-22.
  35. ^ «Science Links Japan | Two Unique Aftercataracts Requiring Surgical Removal». Sciencelinks.jp. 2009-03-18. Archived from the original on 2011-02-17. Retrieved 2010-08-22.
  36. ^ «Dictionary entry for monimolimnion, a word that, at 13 letters, is longer than any of the words linked in the source above». Archived from the original on 2009-09-09. Retrieved 2009-08-15.
  37. ^ «Word Records». Fun-with-words.com. Archived from the original on 2012-08-26. Retrieved 2012-08-13.
  38. ^ «Typewriter Words». Wordnik.com. Archived from the original on 2011-07-17. Retrieved 2011-01-15.
  39. ^ «The Dvorak Keyboard and You». Theworldofstuff.com. Archived from the original on 2010-08-20. Retrieved 2010-08-22.

External links

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  • A Collection of Word Oddities and Trivia – Long words
    • Long words (chemical names)
    • Long words (place names)
  • What is the longest English word?, AskOxford.com «Ask the Experts»
  • What is the Longest Word?, Fun-With-Words.com
  • Full chemical name of titin.
  • Taxonomy of Wordplay
Источник

Ready to take your Scrabble skills to the next level? This list of the longest words in the English language could score you major points on your next game — if you can remember how to spell them.

Some of the words that qualify for the title take hours to pronounce, like the 189,819-letter word for the  protein Titin. Additionally, many of the longest words are medical terms, so we have excluded some of them to allow for more variety. The end result is a list of fascinatingly lengthy words that will make your vocabulary downright sesquipedalian.

Antidisestablishmentarianism

Part of Speech: noun

Definition: opposition to the disestablishment of the Church of England

Origins: While the word originated in 19th century Britain, it is now used to refer to any opposition to a government withdrawing support from a religious organization. Though rarely used in casual conversation, the word was featured in the Duke Ellington song, “You’re Just an Old Antidisestablishmentarianist.”  

Floccinaucinihilipilification

Part of Speech: noun

Definition: the act of defining or estimating something as worthless

Origins: This word stems from the combination of four Latin words, all of which signify that something has little value: flocci, nauci, nihili, pilifi. This style of word creation was popular in Britain in the 1700s. 

Pneumonoultramicroscopicsilicovolcanoconiosis

Part of Speech: noun

Definition: an invented word said to mean a lung disease caused by inhaling a fine dust

Origins: This word emerged in the late 1930s, and was said to be invented by Everett K. Smith, president of the National Puzzlers’ League, in an imitation of very long medical terms. It is not found in real medical usage.

Pseudopseudohypoparathyroidism

Part of Speech: noun

Definition: an inherited disorder similar to pseudohypoparathyroidism

Origins: This genetic disorder causes “short stature, round face and short hand bones,” according to the National Institutes of Health. Despite having a similar name, it is not the same as pseudohypoparathyroidism.

Psychoneuroendocrinological

Part of Speech: adjective

Definition: of or related to to the branch of science concerned with the relationships between psychology, the nervous system, and the endocrine system 

Origins: This term was first seen in the 1970s in Journal of Neurological Science, a medical journal. 

Sesquipedalian

Part of Speech: adjective

Definition: having many syllables or characterized by the use of long words

Origins: The Roman poet Horace used this term to caution young poets against relying on words that used a large number of letters. It was adopted in the 17th century by poets to ridicule their peers who used lengthy words.

Hippopotomonstrosesquippedaliophobia

Part of Speech: noun

Definition: fear of long words

Fun Fact: This word is most often used in humorous contexts. It is an extension of the word sesquipedalophobia, which has the same meaning and is more often used in a formal context.  

Incomprehensibilities

Part of Speech: noun

Definition: things that are impossible to understand or comprehend

Fun Fact: In the 1990s, this word was named the longest word in common usage.

Uncopyrightable

Part of Speech: adjective

Definition: not able or allowed to be protected by copyright

Fun Fact: This word is one of the longest isograms (a word that does not repeat letters) in the English language.

Dermatoglyphics

Part of Speech: noun

Definition: the scientific study of hands, including fingerprints, lines, mounts, and shapes

Fun Fact: Unlike palmistry, this study is based in science and is often used in criminology as a way to identify both perpetrators and victims.

Euouae

Part of Speech: noun

Definition: a type of cadence in medieval music

Fun Fact: While this word might not look as impressive as others on this list, it’s the longest word in the English language to be composed entirely of vowels. (It’s also the word with the longest string of vowels.)

Psychophysicotherapeutics

Part of Speech: noun

Definition: a therapeutic approach that integrates both the mind and body

Fun Fact: While the Oxford Dictionary does not provide an official definition of this word, it is included on their list of the longest words in the English language. 

Otorhinolaryngological

Part of Speech: adjective

Definition: of or relating to the medical specialization involving the ear, nose, and throat

Fun Fact: This medical specialization is more commonly known by its acronym, ENT. 

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Titin is the largest protein yet discovered. Its full chemical name is 189,819 letters long and, depending on how you define a “word” is the longest word in the English language. It is not included in the dictionary – few lexicographers accept chemical formulae as words. But when people go round Googling “What’s the longest word in the English language?”, they don’t really care if it’s a “real” word or not.

So, I present to you the full chemical name of the protein Titin (it’s a long one):

Methionyl­threonyl­threonyl­glutaminyl­alanyl­prolyl­threonyl­phenylalanyl­threonyl­glutaminyl­prolyl­leucyl­glutaminyl­seryl­valyl­valyl­valyl­leucyl­glutamyl­glycyl­seryl­threonyl­alanyl­threonyl­phenylalanyl­glutamyl­alanyl­histidyl­isoleucyl­seryl­glycyl­phenylalanyl­prolyl­valyl­prolyl­glutamyl­valyl­seryl­tryptophyl­phenylalanyl­arginyl­aspartyl­glycyl­glutaminyl­valyl­isoleucyl­seryl­threonyl­seryl­threonyl­leucyl­prolyl­glycyl­valyl­glutaminyl­isoleucyl­seryl­phenylalanyl­seryl­aspartyl­glycyl­arginyl­alanyl­lysyl­leucyl­threonyl­isoleucyl­prolyl­alanyl­valyl­threonyl­lysyl­alanyl­asparaginyl­seryl­glycyl­arginyl­tyrosyl­seryl­leucyl­lysyl­alanyl­threonyl­asparaginyl­glycyl­seryl­glycyl­glutaminyl­alanyl­threonyl­seryl­threonyl­alanyl­glutamyl­leucyl­leucyl­valyl­lysyl­alanyl­glutamyl­threonyl­alanyl­prolyl­prolyl­asparaginyl­phenylalanyl­valyl­glutaminyl­arginyl­leucyl­glutaminyl­seryl­methionyl­threonyl­valyl­arginyl­glutaminyl­glycyl­seryl­glutaminyl­valyl­arginyl­leucyl­glutaminyl­valyl­arginyl­valyl­threonyl­glycyl­isoleucyl­prolyl­threonyl­prolyl­valyl­valyl­lysyl­phenylalanyl­tyrosyl­arginyl­aspartyl­glycyl­alanyl­glutamyl­isoleucyl­glutaminyl­seryl­seryl­leucyl­aspartyl­phenylalanyl­glutaminyl­isoleucyl­seryl­glutaminyl­glutamyl­glycyl­aspartyl­leucyl­tyrosyl­seryl­leucyl­leucyl­isoleucyl­alanyl­glutamyl­alanyl­tyrosyl­prolyl­glutamyl­aspartyl­seryl­glycyl­threonyl­tyrosyl­seryl­valyl­asparaginyl­alanyl­threonyl­asparaginyl­seryl­valyl­glycyl­arginyl­alanyl­threonyl­seryl­threonyl­alanyl­glutamyl­leucyl­leucyl­valyl­glutaminyl­glycyl­glutamyl­glutamyl­glutamyl­valyl­prolyl­alanyl­lysyl­lysyl­threonyl­lysyl­threonyl­isoleucyl­valyl­seryl­threonyl­alanyl­glutaminyl­isoleucyl­seryl­glutamyl­seryl­arginyl­glutaminyl­threonyl­arginyl­isoleucyl­glutamyl­lysyl­lysyl­isoleucyl­glutamyl­alanyl­histidyl­phenylalanyl­aspartyl­alanyl­arginyl­seryl­isoleucyl­alanyl­threonyl­valyl­glutamyl­methionyl­valyl­isoleucyl­aspartyl­glycyl­alanyl­alanyl­glycyl­glutaminyl­glutaminyl­leucyl­prolyl­histidyl­lysyl­threonyl­prolyl­prolyl­arginyl­isoleucyl­prolyl­prolyl­lysyl­prolyl­lysyl­seryl­arginyl­seryl­prolyl­threonyl­prolyl­prolyl­seryl­isoleucyl­alanyl­alanyl­lysyl­alanyl­glutaminyl­leucyl­alanyl­arginyl­glutaminyl­glutaminyl­seryl­prolyl­seryl­prolyl­isoleucyl­arginyl­histidyl­seryl­prolyl­seryl­prolyl­valyl­arginyl­histidyl­valyl­arginyl­alanyl­prolyl­threonyl­prolyl­seryl­prolyl­valyl­arginyl­seryl­valyl­seryl­prolyl­alanyl­alanyl­arginyl­isoleucyl­seryl­threonyl­seryl­prolyl­isoleucyl­arginyl­seryl­valyl­arginyl­seryl­prolyl­leucyl­leucyl­methionyl­arginyl­lysyl­threonyl­glutaminyl­alanyl­seryl­threonyl­valyl­alanyl­threonyl­glycyl­prolyl­glutamyl­valyl­prolyl­prolyl­prolyl­tryptophyl­lysyl­glutaminyl­glutamyl­glycyl­tyrosyl­valyl­alanyl­seryl­seryl­seryl­glutamyl­alanyl­glutamyl­methionyl­arginyl­glutamyl­threonyl­threonyl­leucyl­threonyl­threonyl­seryl­threonyl­glutaminyl­isoleucyl­arginyl­threonyl­glutamyl­glutamyl­arginyl­tryptophyl­glutamyl­glycyl­arginyl­tyrosyl­glycyl­valyl­glutaminyl­glutamyl­glutaminyl­valyl­threonyl­isoleucyl­seryl­glycyl­alanyl­alanyl­glycyl­alanyl­alanyl­alanyl­seryl­valyl­seryl­alanyl­seryl­alanyl­seryl­tyrosyl­alanyl­alanyl­glutamyl­alanyl­valyl­alanyl­threonyl­glycyl­alanyl­lysyl­glutamyl­valyl­lysyl­glutaminyl­aspartyl­alanyl­aspartyl­lysyl­seryl­alanyl­alanyl­valyl­alanyl­threonyl­valyl­valyl­alanyl­alanyl­valyl­aspartyl­methionyl­alanyl­arginyl­valyl­arginyl­glutamyl­prolyl­valyl­isoleucyl­seryl­alanyl­valyl­glutamyl­glutaminyl­threonyl­alanyl­glutaminyl­arginyl­threonyl­threonyl­threonyl­threonyl­alanyl­valyl­histidyl­isoleucyl­glutaminyl­prolyl­alanyl­glutaminyl­glutamyl­glutaminyl­valyl­arginyl­lysyl­glutamyl­alanyl­glutamyl­lysyl­threonyl­alanyl­valyl­threonyl­lysyl­valyl­valyl­valyl­alanyl­alanyl­aspartyl­lysyl­alanyl­lysyl­glutamyl­glutaminyl­glutamyl­leucyl­lysyl­seryl­arginyl­threonyl­lysyl­glutamyl­valyl­isoleucyl­threonyl­threonyl­lysyl­glutaminyl­glutamyl­glutaminyl­methionyl­histidyl­valyl­threonyl­histidyl­glutamyl­glutaminyl­isoleucyl­arginyl­lysyl­glutamyl­threonyl­glutamyl­lysyl­threonyl­phenylalanyl­valyl­prolyl­lysyl­valyl­valyl­isoleucyl­seryl­alanyl­alanyl­lysyl­alanyl­lysyl­glutamyl­glutaminyl­glutamyl­threonyl­arginyl­isoleucyl­seryl­glutamyl­glutamyl­isoleucyl­threonyl­lysyl­lysyl­glutaminyl­lysyl­glutaminyl­valyl­threonyl­glutaminyl­glutamyl­alanyl­isoleucyl­arginyl­glutaminyl­glutamyl­threonyl­glutamyl­isoleucyl­threonyl­alanyl­alanyl­seryl­methionyl­valyl­valyl­valyl­alanyl­threonyl­alanyl­lysyl­seryl­threonyl­lysyl­leucyl­glutamyl­threonyl­valyl­prolyl­glycyl­alanyl­glutaminyl­glutamyl­glutamyl­threonyl­threonyl­threonyl­glutaminyl­glutaminyl­aspartyl­glutaminyl­methionyl­histidyl­leucyl­seryl­tyrosyl­glutamyl­lysyl­isoleucyl­methionyl­lysyl­glutamyl­threonyl­arginyl­lysyl­threonyl­valyl­valyl­prolyl­lysyl­valyl­isoleucyl­valyl­alanyl­threonyl­prolyl­lysyl­valyl­lysyl­glutamyl­glutaminyl­aspartyl­leucyl­valyl­seryl­arginyl­glycyl­arginyl­glutamyl­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yl­isoleucyl­leucyl­glycyl­tyrosyl­isoleucyl­valyl­glutamyl­tyrosyl­glutaminyl­lysyl­valyl­glycyl­aspartyl­glutamyl­glutamyl­tryptophyl­arginyl­arginyl­alanyl­asparaginyl­histidyl­threonyl­prolyl­glutamyl­seryl­cysteinyl­prolyl­glutamyl­threonyl­lysyl­tyrosyl­lysyl­valyl­threonyl­glycyl­leucyl­arginyl­aspartyl­glycyl­glutaminyl­threonyl­tyrosyl­lysyl­phenylalanyl­arginyl­valyl­leucyl­alanyl­valyl­asparaginyl­alanyl­alanyl­glycyl­glutamyl­seryl­aspartyl­prolyl­alanyl­histidyl­valyl­prolyl­glutamyl­prolyl­valyl­leucyl­valyl­lysyl­aspartyl­arginyl­leucyl­glutamyl­prolyl­prolyl­glutamyl­leucyl­isoleucyl­leucyl­aspartyl­alanyl­asparaginyl­methionyl­alanyl­arginyl­glutamyl­glutaminyl­histidyl­isoleucyl­lysyl­valyl­glycyl­aspartyl­threonyl­leucyl­arginyl­leucyl­seryl­alanyl­isoleucyl­isoleucyl­lysyl­glycyl­valyl­prolyl­phenylalanyl­prolyl­lysyl­valyl­threonyl­tryptophyl­lysyl­lysyl­glutamyl­aspartyl­arginyl­aspartyl­alanyl­prolyl­threonyl­lysyl­alanyl­arginyl­isoleucyl­aspartyl­valyl­threonyl­prolyl­valyl­glycyl­seryl­lysyl­leucyl­glutamyl­isoleucyl­arginyl­asparaginyl­alanyl­alanyl­histidyl­glutamyl­aspartyl­glycyl­glycyl­isoleucyl­tyrosyl­seryl­leucyl­threonyl­valyl­glutamyl­asparaginyl­prolyl­alanyl­glycyl­seryl­lysyl­threonyl­valyl­seryl­valyl­lysyl­valyl­leucyl­valyl­leucyl­aspartyl­lysyl­prolyl­glycyl­prolyl­prolyl­arginyl­aspartyl­leucyl­glutamyl­valyl­seryl­glutamyl­isoleucyl­arginyl­lysyl­aspartyl­seryl­cysteinyl­tyrosyl­leucyl­threonyl­tryptophyl­lysyl­glutamyl­prolyl­leucyl­aspartyl­aspartyl­glycyl­glycyl­seryl­valyl­isoleucyl­threonyl­asparaginyl­tyrosyl­valyl­valyl­glutamyl­arginyl­arginyl­aspartyl­valyl­alanyl­seryl­alanyl­glutaminyl­tryptophyl­seryl­prolyl­leucyl­seryl­alanyl­threonyl­seryl­lysyl­lysyl­lysyl­seryl­histidyl­phenylalanyl­alanyl­lysyl­histidyl­leucyl­asparaginyl­glutamyl­glycyl­asparaginyl­glutaminyl­tyrosyl­leucyl­phenylalanyl­arginyl­valyl­alanyl­alanyl­glutamyl­asparaginyl­glutaminyl­tyrosyl­glycyl­arginyl­glycyl­prolyl­phenylalanyl­valyl­glutamyl­threonyl­prolyl­lysyl­prolyl­isoleucyl­lysyl­alanyl­leucyl­aspartyl­prolyl­leucyl­histidyl­prolyl­prolyl­glycyl­prolyl­prolyl­lysyl­aspartyl­leucyl­histidyl­histidyl­valyl­aspartyl­valyl­aspartyl­lysyl­threonyl­glutamyl­valyl­seryl­leucyl­valyl­tryptophyl­asparaginyl­lysyl­prolyl­aspartyl­arginyl­aspartyl­glycyl­glycyl­seryl­prolyl­isoleucyl­threonyl­glycyl­tyrosyl­leucyl­valyl­glutamyl­tyrosyl­glutaminyl­glutamyl­glutamyl­glycyl­threonyl­glutaminyl­aspartyl­tryptophyl­isoleucyl­lysyl­phenylalanyl­lysyl­threonyl­valyl­threonyl­asparaginyl­leucyl­glutamyl­cysteinyl­valyl­valyl­threonyl­glycyl­leucyl­glutaminyl­glutaminyl­glycyl­lysyl­threonyl­tyrosyl­arginyl­phenylalanyl­arginyl­valyl­lysyl­alanyl­glutamyl­asparaginyl­isoleucyl­valyl­glycyl­leucyl­glycyl­leucyl­prolyl­aspartyl­threonyl­threonyl­isoleucyl­prolyl­isoleucyl­glutamyl­cysteinyl­glutaminyl­glutamyl­lysyl­leucyl­valyl­prolyl­prolyl­seryl­valyl­glutamyl­leucyl­aspartyl­valyl­lysyl­leucyl­isoleucyl­glutamyl­glycyl­leucyl­valyl­valyl­lysyl­alanyl­glycyl­threonyl­threonyl­valyl­arginyl­phenylalanyl­prolyl­alanyl­isoleucyl­isoleucyl­arginyl­glycyl­valyl­prolyl­valyl­prolyl­threonyl­alanyl­lysyl­tryptophyl­threonyl­threonyl­aspartyl­glycyl­seryl­glutamyl­isoleucyl­lysyl­threonyl­aspartyl­glutamyl­histidyl­tyrosyl­threonyl­valyl­glutamyl­threonyl­aspartyl­asparaginyl­phenylalanyl­seryl­seryl­valyl­leucyl­threonyl­isoleucyl­lysyl­asparaginyl­cysteinyl­leucyl­arginyl­arginyl­aspartyl­threonyl­glycyl­glutamyl­tyrosyl­glutaminyl­isoleucyl­threonyl­valyl­seryl­asparaginyl­alanyl­alanyl­glycyl­seryl­lysyl­threonyl­valyl­alanyl­valyl­histidyl­leucyl­threonyl­valyl­leucyl­aspartyl­valyl­prolyl­glycyl­prolyl­prolyl­threonyl­glycyl­prolyl­isoleucyl­asparaginyl­isoleucyl­leucyl­aspartyl­valyl­threonyl­prolyl­glutamyl­histidyl­methionyl­threonyl­isoleucyl­seryl­tryptophyl­glutaminyl­prolyl­prolyl­lysyl­aspartyl­aspartyl­glycyl­glycyl­seryl­prolyl­valyl­isoleucyl­asparaginyl­tyrosyl­isoleucyl­valyl­glutamyl­lysyl­glutaminyl­aspartyl­threonyl­arginyl­lysyl­aspartyl­threonyl­tryptophyl­glycyl­valyl­valyl­seryl­seryl­glycyl­seryl­seryl­lysyl­threonyl­lysyl­leucyl­lysyl­isoleucyl­prolyl­histidyl­leucyl­glutaminyl­lysyl­glycyl­cysteinyl­glutamyl­tyrosyl­valyl­phenylalanyl­arginyl­valyl­arginyl­alanyl­glutamyl­asparaginyl­lysyl­isoleucyl­glycyl­valyl­glycyl­prolyl­prolyl­leucyl­aspartyl­seryl­threonyl­prolyl­threonyl­valyl­alanyl­lysyl­histidyl­lysyl­phenylalanyl­seryl­prolyl­prolyl­seryl­prolyl­prolyl­glycyl­lysyl­prolyl­valyl­valyl­threonyl­aspartyl­isoleucyl­threonyl­glutamyl­asparaginyl­alanyl­alanyl­threonyl­valyl­seryl­tryptophyl­threonyl­leucyl­prolyl­lysyl­seryl­aspartyl­glycyl­glycyl­seryl­prolyl­isoleucyl­threonyl­glycyl­tyrosyl­tyrosyl­methionyl­glutamyl­arginyl­arginyl­glutamyl­valyl­threonyl­glycyl­lysyl­tryptophyl­valyl­arginyl­valyl­asparaginyl­lysyl­threonyl­prolyl­isoleucyl­alanyl­aspartyl­leucyl­lysyl­phenylalanyl­arginyl­valyl­threonyl­glycyl­leucyl­tyrosyl­glutamyl­glycyl­asparaginyl­th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­seryl­prolyl­glutamyl­prolyl­seryl­histidyl­prolyl­lysyl­alanyl­valyl­seryl­prolyl­threonyl­glutamyl­threonyl­lysyl­prolyl­threonyl­prolyl­threonyl­glutamyl­lysyl­valyl­glutaminyl­histidyl­leucyl­prolyl­valyl­seryl­alanyl­prolyl­prolyl­lysyl­isoleucyl­threonyl­glutaminyl­phenylalanyl­leucyl­lysyl­alanyl­glutamyl­alanyl­seryl­lysyl­glutamyl­isoleucyl­alanyl­lysyl­leucyl­threonyl­cysteinyl­valyl­valyl­glutamyl­seryl­seryl­valyl­leucyl­arginyl­alanyl­lysyl­glutamyl­valyl­threonyl­tryptophyl­tyrosyl­lysyl­aspartyl­glycyl­lysyl­lysyl­leucyl­lysyl­glutamyl­asparaginyl­glycyl­histidyl­phenylalanyl­glutaminyl­phenylalanyl­histidyl­tyrosyl­seryl­alanyl­aspartyl­glycyl­threonyl­tyrosyl­glutamyl­leucyl­lysyl­isoleucyl­asparaginyl­asparaginyl­leucyl­threonyl­glutamyl­seryl­aspartyl­glutaminyl­glycyl­glutamyl­tyrosyl­valyl­cysteinyl­glutamyl­isoleucyl­seryl­glycyl­glutamyl­glycyl­glycyl­threonyl­seryl­lysyl­threonyl­asparaginyl­leucyl­glutaminyl­phenylalanyl­methionyl­glycyl­glutaminyl­alanyl­phenylalanyl­lysyl­seryl­isoleucyl­histidyl­glutamyl­lysyl­valyl­seryl­lysyl­isoleucyl­seryl­glutamyl­threonyl­lysyl­lysyl­seryl­aspartyl­glutaminyl­lysyl­threonyl­threonyl­glutamyl­seryl­threonyl­valyl­threonyl­arginyl­lysyl­threonyl­glutamyl­prolyl­lysyl­alanyl­prolyl­glutamyl­prolyl­isoleucyl­seryl­seryl­lysyl­prolyl­valyl­isoleucyl­valyl­threonyl­glycyl­leucyl­glutaminyl­aspartyl­threonyl­threonyl­valyl­seryl­seryl­aspartyl­seryl­valyl­alanyl­lysyl­phenylalanyl­alanyl­valyl­lysyl­alanyl­threonyl­glycyl­glutamyl­prolyl­arginyl­prolyl­threonyl­alanyl­isoleucyl­tryptophyl­threonyl­lysyl­aspartyl­glycyl­lysyl­alanyl­isoleucyl­threonyl­glutaminyl­glycyl­glycyl­lysyl­tyrosyl­lysyl­leucyl­seryl­glutamyl­aspartyl­lysyl­glycyl­glycyl­phenylalanyl­phenylalanyl­leucyl­glutamyl­isoleucyl­histidyl­lysyl­threonyl­aspartyl­threonyl­seryl­aspartyl­seryl­glycyl­leucyl­tyrosyl­threonyl­cysteinyl­threonyl­valyl­lysyl­asparaginyl­seryl­alanyl­glycyl­seryl­valyl­seryl­seryl­seryl­cysteinyl­lysyl­leucyl­threonyl­isoleucyl­lysyl­alanyl­isoleucyl­lysyl­aspartyl­threonyl­glutamyl­alanyl­glutaminyl­lysyl­valyl­seryl­threonyl­glutaminyl­lysyl­threonyl­seryl­glutamyl­isoleucyl­threonyl­prolyl­glutaminyl­lysyl­lysyl­alanyl­valyl­valyl­glutaminyl­glutamyl­glutamyl­isoleucyl­seryl­glutaminyl­lysyl­alanyl­leucyl­arginyl­seryl­glutamyl­glutamyl­isoleucyl­lysyl­methionyl­seryl­glutamyl­alanyl­lysyl­seryl­glutaminyl­glutamyl­lysyl­leucyl­alanyl­leucyl­lysyl­glutamyl­glutamyl­alanyl­seryl­lysyl­valyl­leucyl­isoleucyl­seryl­glutamyl­glutamyl­valyl­lysyl­lysyl­seryl­alanyl­alanyl­threonyl­seryl­leucyl­glutamyl­lysyl­seryl­isoleucyl­valyl­histidyl­glutamyl­glutamyl­isoleucyl­threonyl­lysyl­threonyl­seryl­glutaminyl­alanyl­seryl­glutamyl­glutamyl­valyl­arginyl­threonyl­histidyl­alanyl­glutamyl­isoleucyl­lysyl­alanyl­phenylalanyl­seryl­threonyl­glutaminyl­methionyl­seryl­isoleucyl­asparaginyl­glutamyl­glycyl­glutaminyl­arginyl­leucyl­valyl­leucyl­lysyl­alanyl­asparaginyl­isoleucyl­alanyl­glycyl­alanyl­threonyl­aspartyl­valyl­lysyl­tryptophyl­valyl­leucyl­asparaginyl­glycyl­valyl­glutamyl­leucyl­threonyl­asparaginyl­seryl­glutamyl­glutamyl­tyrosyl­arginyl­tyrosyl­glycyl­valyl­seryl­glycyl­seryl­aspartyl­glutaminyl­threonyl­leucyl­threonyl­isoleucyl­lysyl­glutaminyl­alanyl­seryl­histidyl­arginyl­aspartyl­glutamyl­glycyl­isoleucyl­leucyl­threonyl­cysteinyl­isoleucyl­seryl­lysyl­threonyl­lysyl­glutamyl­glycyl­isoleucyl­valyl­lysyl­cysteinyl­glutaminyl­tyrosyl­aspartyl­leucyl­threonyl­leucyl­seryl­lysyl­glutamyl­leucyl­seryl­aspartyl­alanyl­prolyl­alanyl­phenylalanyl­isoleucyl­seryl­glutaminyl­prolyl­arginyl­seryl­glutaminyl­asparaginyl­isoleucyl­asparaginyl­glutamyl­glycyl­glutaminyl­asparaginyl­valyl­leucyl­phenylalanyl­threonyl­cysteinyl­glutamyl­isoleucyl­seryl­glycyl­glutamyl­prolyl­seryl­prolyl­glutamyl­isoleucyl­glutamyl­tryptophyl­phenylalanyl­lysyl­asparaginyl­asparaginyl­leucyl­prolyl­isoleucyl­seryl­isoleucyl­seryl­seryl­asparaginyl­valyl­seryl­isoleucyl­seryl­arginyl­seryl­arginyl­asparaginyl­valyl­tyrosyl­seryl­leucyl­glutamyl­isoleucyl­arginyl­asparaginyl­alanyl­seryl­valyl­seryl­aspartyl­seryl­glycyl­lysyl­tyrosyl­threonyl­isoleucyl­lysyl­alanyl­lysyl­asparaginyl­phenylalanyl­arginyl­glycyl­glutaminyl­cysteinyl­seryl­alanyl­threonyl­alanyl­seryl­leucyl­methionyl­valyl­leucyl­prolyl­leucyl­valyl­glutamyl­glutamyl­prolyl­seryl­arginyl­glutamyl­valyl­valyl­leucyl­arginyl­threonyl­seryl­glycyl­aspartyl­threonyl­seryl­leucyl­glutaminyl­glycyl­seryl­phenylalanyl­seryl­seryl­glutaminyl­seryl­valyl­glutaminyl­methionyl­seryl­alanyl­seryl­lysyl­glutaminyl­glutamyl­alanyl­seryl­phenylalanyl­seryl­seryl­phenylalanyl­seryl­seryl­seryl­seryl­alanyl­seryl­seryl­methionyl­threonyl­glutamyl­methionyl­lysyl­phenylalanyl­alanyl­seryl­methionyl­seryl­alanyl­glutaminyl­seryl­methionyl­seryl­seryl­methionyl­glutaminyl­glutamyl­seryl­phenylalanyl­valyl­glutamyl­methionyl­seryl­seryl­seryl­seryl­phenylalanyl­methionyl­glycyl­isoleucyl­seryl­asparaginyl­methionyl­threonyl­glutaminyl­leucyl­glutamyl­seryl­seryl­threonyl­seryl­lysyl­methionyl­leucyl­lysyl­alanyl­glycyl­isoleucyl­arginyl­glycyl­isoleucyl­prolyl­prolyl­lysyl­isoleucyl­glutamyl­alanyl­leucyl­prolyl­seryl­aspartyl­isoleucyl­seryl­isoleucyl­aspartyl­glutamyl­glycyl­lysyl­valyl­leucyl­threonyl­valyl­alanyl­cysteinyl­alanyl­phenylalanyl­threonyl­glycyl­glutamyl­prolyl­threonyl­prolyl­glutamyl­valyl­threonyl­tryptophyl­seryl­cysteinyl­glycyl­glycyl­arginyl­lysyl­isoleucyl­histidyl­seryl­glutaminyl­glutamyl­glutaminyl­glycyl­arginyl­phenylalanyl­histidyl­isoleucyl­glutamyl­asparaginyl­threonyl­aspartyl­aspartyl­leucyl­threonyl­threonyl­leucyl­isoleucyl­isoleucyl­methionyl­aspartyl­valyl­glutaminyl­lysyl­glutaminyl­aspartyl­glycyl­glycyl­leucyl­tyrosyl­threonyl­leucyl­seryl­leucyl­glycyl­asparaginyl­glutamyl­phenylalanyl­glycyl­seryl­aspartyl­seryl­alanyl­threonyl­valyl­asparaginyl­isoleucyl­histidyl­isoleucyl­arginyl­seryl­isoleucine.

Источник

A man has been recorded spending more than three hours to pronounce what is supposedly the longest word in the English language.

‘Methionylthreonylthreonylglutaminylarginyl…isoleucine’ is the chemical name of ‘titin’ (also known as ‘connectin’) — the largest known protein. It has 189,819 letters.

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However, in the past it has been disputed whether or not it is a real word — proteins are named after the chemicals involved in making them.

The longest word in the Oxford English Dictionary is ‘pneumonoultramicroscopicsilicovolcanoconiosis’ at 45 letters long.

‘Supercalifragilisticexpialidocious’, made famous by Mary Poppins, is 34 letters long.

> ‘Amazeballs’, ‘mummy porn’, ‘frenemy’ make Collins online dictionary

> OMG, LOL enter Oxford English Dictionary

Below is the full 189,819-lettered word for ‘titin’:

Methionylthreonylthreonylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylglutaminylleuc yllysylglutamylarginyllysylglutamylglycylalanylphenylalanylvalylprolylphenylalanylvalylthreonylleucylgl ycylaspartylprolylglycylisoleucylglutamylglutaminylserylleucyllysylisoleucylaspartylthreonylleucylisoleu cylglutamylalanylglycylalanylaspartylalanylleucylglutamylleucylglycylisoleucylprolylphenylalanylseryla spartylprolylleucylalanylaspartylglycylprolylthreonylisoleucylglutaminylasparaginylalanylthreonylleucyl arginylalanylphenylalanylalanylalanylglycylvalylthreonylprolylalanylglutaminylcysteinylphenylalanylglu tamylmethionylleucylalanylleucylisoleucylarginylglutaminyllysylhistidylprolylthreonylisoleucylprolylisol eucylglycylleucylleucylmethionyltyrosylalanylasparaginylleucylvalylphenylalanylasparaginyllysylglycyli soleucylaspartylglutamylphenylalanyltyrosylalanylglutaminylcysteinylglutamyllysylvalylglycylvalylaspa rtylserylvalylleucylvalylalanylaspartylvalylprolylvalylglutaminylglutamylserylalanylprolylphenylalanylarg inylglutaminylalanylalanylleucylarginylhistidylasparaginylvalylalanylprolylisoleucylphenylalanylisoleuc ylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalany lseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylgly cylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanylly sylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycy lisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylala nylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhist idylasparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylal anylvalylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreo nylserylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartyl prolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylgl utaminylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylval ylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminy lserylthreonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylaspar aginylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalany laspartylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminy lglutaminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylv 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ysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucy lserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycyl alanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylaspa raginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalyl glutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylp rolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisol eucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminyl phenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutam inylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthr 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ucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylp rolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglu taminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutam ylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenylalanyl glutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutamin ylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylar ginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleu cylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreonylar ginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylserylpr 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lylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucyl serylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucy lglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylproly lmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylgluta minylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamyll eucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenylalanylgl utaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminyl phenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylargi nylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleucy laspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreonylargin ylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylserylprolylt hreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartylaspartylal anylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalylasparagin ylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylthreonylp henylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylalanyltitinme thionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylprolylaspar tylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylgly cyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparagin ylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosyl asparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylas partylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylseryl glycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglut amylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmet hionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminyl phenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucyl leucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenylalanylglutami nylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylpheny lalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylarginylphe nylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleucylaspar tylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreonylarginylaspa raginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylserylprolylthreon ylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartylaspartylalanylth reonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalylasparaginylgluta mylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylthreonylphenyla lanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylalanyltitinmethionyl glutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylprolylaspartylalan ylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyro sylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginylargin ylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylaspar aginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartyl glutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycy lserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamyl prolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethion yllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphen ylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleuc 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lasparaginylleucylvalylphenylalanylasparaginyllysylglycylisoleucylaspartylglutamylphenylalanyltyrosyl alanylglutaminylcysteinylglutamyllysylvalylglycylvalylaspartylserylvalylleucylvalylalanylaspartylvalylpro lylvalylglutaminylglutamylserylalanylprolylphenylalanylarginylglutaminylalanylalanylleucylarginylhistidy lasparaginylvalylalanylprolylisoleucylphenylalanylisoleucylcysteinylprolylprolylaspartylalanylaspartylas partylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonylt yrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginylarginylalanylalan ylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanyl alanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylv alyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanyli soleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylprolylglutam yllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanyl alanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenylalanylvaly lphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleucylasparagin ylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenylalanylglutaminylthreonylgluta minylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphenylalanylserylgluta minylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylarginylphenylalanylprolylgl ycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleucylaspartylprolylleucyliso leucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreonylarginylasparaginylarginyliso 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ylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalylly sylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisole ucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylprolylglutamyllys ylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanylalan ylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenylalanylvalylphe nylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleucylasparaginylval ylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenylalanylglutaminylthreonylglutamin ylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphenylalanylserylglutamin ylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylarginylphenylalanylprolylglycy laspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleucylaspartylprolylleucylisoleu cylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreonylarginylasparaginylarginylisoleu cylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylserylprolylthreonylthreonylalanylglut amylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartylaspartylalanylthreonylvalylalanyliso leucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalylasparaginylglutamylleucylvalylargin ylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylthreonylphenylalanylglutamylseryl methionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylalanyltitinmethionylglutaminylarginylt yrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylprolylaspartylalanylaspartylaspartyl aspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosyl 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utaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphenylalanylserylglutaminylv alyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylarginylphenylalanylprolylglycylas partylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleucylaspartylprolylleucylisoleucylt hreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreonylarginylasparaginylarginylisoleucyli soleucylglutamylvalylglutamylasparaginylglutaminylglutaminylserylprolylthreonylthreonylalanylglutam ylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartylaspartylalanylthreonylvalylalanylisoleu cylarginylserylalanylasparaginylisoleucylasparaginylleucylvalylasparaginylglutamylleucylvalylarginylgl ycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylthreonylphenylalanylglutamylserylmet hionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylalanyltitinmethionylglutaminylarginyltyros ylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylprolylaspartylalanylaspartylaspartylasp artylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleu cylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginylarginylalanylalanylleucylp rolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanylalanylprol ylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalyllysylal anylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisoleucylv alyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylprolylglutamyllysylmet hionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanylalanylthre onylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenylalanylvalylphenylala 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ylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylthreonylphenylalanylglutamylserylmethi onylserylglycylleucylvalyltryptophylthreonylserylalanylprolylalanyltitinmethionylglutaminylarginyltyrosyl glutamylserylleucylphenylalanylalanylmethionylthreonylthreonylglutaminylarginyltyrosylglutamylseryll eucylphenylalanylalanylglutaminylleucyllysylglutamylarginyllysylglutamylglycylalanylphenylalanylvalyl prolylphenylalanylvalylthreonylleucylglycylaspartylprolylglycylisoleucylglutamylglutaminylserylleucylly sylisoleucylaspartylthreonylleucylisoleucylglutamylalanylglycylalanylaspartylalanylleucylglutamylleucy lglycylisoleucylprolylphenylalanylserylaspartylprolylleucylalanylaspartylglycylprolylthreonylisoleucylglu taminylasparaginylalanylthreonylleucylarginylalanylphenylalanylalanylalanylglycylvalylthreonylprolylal anylglutaminylcysteinylphenylalanylglutamylmethionylleucylalanylleucylisoleucylarginylglutaminyllysyl 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ylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysyli soleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylprolylglutamyllysylmethionylle ucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanylalanylthreonylargi nylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenylalanylvalylphenylalanylleuc ylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthr eonylserylserylleucylglycylasparaginylglutaminylphenylalanylglutaminylthreonylglutaminylglutaminylal anylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptop hyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylarginylphenylalanylprolylglycylaspartylvalyl tyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleucylaspartylprolylleucylisoleucylthreonylal anylleucylleucylglycylthreonylphenylalanylaspartylthreonylarginylasparaginylarginylisoleucylisoleucylg lutamylvalylglutamylasparaginylglutaminylglutaminylserylprolylthreonylthreonylalanylglutamylthreonyll eucylaspartylalanylthreonylarginylarginylvalylaspartylaspartylalanylthreonylvalylalanylisoleucylarginyl serylalanylasparaginylisoleucylasparaginylleucylvalylasparaginylglutamylleucylvalylarginylglycylthreo nylglycylleucyltyrosylasparaginylglutaminylasparaginylthreonylphenylalanylglutamylserylmethionylser ylglycylleucylvalyltryptophylthreonylserylalanylprolylalanyltitinmethionylglutaminylarginyltyrosylglutam ylserylleucylphenylalanylalanylisoleucylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleuc ylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucyl serylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginylarginylalanylalanylleucylprolylleuc ylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolyll eucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalan ylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysyli soleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylprolylglutamyllysylmethionylle ucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanylalanylthreonylargi nylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenylalanylvalylphenylalanylleuc ylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthr eonylserylserylleucylglycylasparaginylglutaminylphenylalanylglutaminylthreonylglutaminylglutaminylal anylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptop 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serylalanylasparaginylisoleucylasparaginylleucylvalylasparaginylglutamylleucylvalylarginylglycylthreo nylglycylleucyltyrosylasparaginylglutaminylasparaginylthreonylphenylalanylglutamylserylmethionylser ylglycylleucylvalyltryptophylthreonylserylalanylprolylalanyltitinmethionylglutaminylarginyltyrosylglutam ylserylleucylphenylalanylalanylmethionylthreonylthreonylglutaminylarginyltyrosylglutamylserylleucylph enylalanylalanylglutaminylleucyllysylglutamylarginyllysylglutamylglycylalanylphenylalanylvalylprolylph enylalanylvalylthreonylleucylglycylaspartylprolylglycylisoleucylglutamylglutaminylserylleucyllysylisoleu cylaspartylthreonylleucylisoleucylglutamylalanylglycylalanylaspartylalanylleucylglutamylleucylglycyliso leucylprolylphenylalanylserylaspartylprolylleucylalanylaspartylglycylprolylthreonylisoleucylglutaminyla sparaginylalanylthreonylleucylarginylalanylphenylalanylalanylalanylglycylvalylthreonylprolylalanylgluta minylcysteinylphenylalanylglutamylmethionylleucylalanylleucylisoleucylarginylglutaminyllysylhistidylpr olylthreonylisoleucylprolylisoleucylglycylleucylleucylmethionyltyrosylalanylasparaginylleucylvalylphen ylalanylasparaginyllysylglycylisoleucylaspartylglutamylphenylalanyltyrosylalanylglutaminylcysteinylglu tamyllysylvalylglycylvalylaspartylserylvalylleucylvalylalanylaspartylvalylprolylvalylglutaminylglutamylse rylalanylprolylphenylalanylarginylglutaminylalanylalanylleucylarginylhistidylasparaginylvalylalanylproly lisoleucylphenylalanylisoleucylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylar ginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylargin ylalanylglycylvalylthreonylglycylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparagi nylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylgluta minylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucyl aspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylis oleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanyl alanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetyls eryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylsery lvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylseryls erylleucylglycylasparaginylglutaminylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylt hreonylthreonylglutaminylvalylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprol ylphenylalanylprolylglutaminylserylthreonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysyl 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asparaginylglutaminylasparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryp tophylthreonylserylalanylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylala nylalanylisoleucylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutamin ylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycyl valylthreonylglycylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleu cylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylph enylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalany lglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutam ylglutaminylhistidylasparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllys 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ylarginylarginylvalylaspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisol eucylasparaginylleucylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylaspar aginylglutaminylasparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophy lthreonylserylalanylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylal anylmethionylthreonylthreonylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylglutaminyll eucyllysylglutamylarginyllysylglutamylglycylalanylphenylalanylvalylprolylphenylalanylvalylthreonylleuc ylglycylaspartylprolylglycylisoleucylglutamylglutaminylserylleucyllysylisoleucylaspartylthreonylleucylis oleucylglutamylalanylglycylalanylaspartylalanylleucylglutamylleucylglycylisoleucylprolylphenylalanylse rylaspartylprolylleucylalanylaspartylglycylprolylthreonylisoleucylglutaminylasparaginylalanylthreonylle ucylarginylalanylphenylalanylalanylalanylglycylvalylthreonylprolylalanylglutaminylcysteinylphenylalany lglutamylmethionylleucylalanylleucylisoleucylarginylglutaminyllysylhistidylprolylthreonylisoleucylprolyli soleucylglycylleucylleucylmethionyltyrosylalanylasparaginylleucylvalylphenylalanylasparaginyllysylgly cylisoleucylaspartylglutamylphenylalanyltyrosylalanylglutaminylcysteinylglutamyllysylvalylglycylvalyla spartylserylvalylleucylvalylalanylaspartylvalylprolylvalylglutaminylglutamylserylalanylprolylphenylalany larginylglutaminylalanylalanylleucylarginylhistidylasparaginylvalylalanylprolylisoleucylphenylalanylisol eucylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylal anylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreony lglycylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalan 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glutaminylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylv alylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutami nylserylthreonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylas paraginylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylal anylaspartylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylgluta minylglutaminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylar ginylvalylaspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylaspar aginylleucylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylgluta minylasparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylse rylalanylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleuc ylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalany lseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylgly cylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanylly sylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycy lisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylala nylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhist idylasparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylal anylvalylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreo nylserylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartyl prolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylgl utaminylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylval ylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminy lserylthreonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylaspar aginylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalany laspartylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminy lglutaminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylv alylaspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginyl leucylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminyl asparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylala nylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcyst einylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylserylt yrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylala nylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleu cyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisole ucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylgl ycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidyla sparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylv alylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylser ylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolyli soleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutami nylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglut aminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylseryl threonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparagin ylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylasp artylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglut aminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalyl aspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleu cylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasp araginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanyl prolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylmethionylthreon ylthreonylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylglutaminylleucyllysylglutamylar ginyllysylglutamylglycylalanylphenylalanylvalylprolylphenylalanylvalylthreonylleucylglycylaspartylprolyl glycylisoleucylglutamylglutaminylserylleucyllysylisoleucylaspartylthreonylleucylisoleucylglutamylalanyl glycylalanylaspartylalanylleucylglutamylleucylglycylisoleucylprolylphenylalanylserylaspartylprolylleucy lalanylaspartylglycylprolylthreonylisoleucylglutaminylasparaginylalanylthreonylleucylarginylalanylphen ylalanylalanylalanylglycylvalylthreonylprolylalanylglutaminylcysteinylphenylalanylglutamylmethionylleu cylalanylleucylisoleucylarginylglutaminyllysylhistidylprolylthreonylisoleucylprolylisoleucylglycylleucylle ucylmethionyltyrosylalanylasparaginylleucylvalylphenylalanylasparaginyllysylglycylisoleucylaspartylgl utamylphenylalanyltyrosylalanylglutaminylcysteinylglutamyllysylvalylglycylvalylaspartylserylvalylleucyl valylalanylaspartylvalylprolylvalylglutaminylglutamylserylalanylprolylphenylalanylarginylglutaminylalan ylalanylleucylarginylhistidylasparaginylvalylalanylprolylisoleucylphenylalanylisoleucylcysteinylprolylpr olylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycyl arginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla spartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalyla sparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylt hreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylala nyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylp rolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycy larginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla spartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalyla sparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylt hreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylala nyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylp rolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycy larginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla 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oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla spartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalyla sparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylt hreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylala nyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylmethionylthreonylthreon ylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylglutaminylleucyllysylglutamylarginyllys ylglutamylglycylalanylphenylalanylvalylprolylphenylalanylvalylthreonylleucylglycylaspartylprolylglycylis oleucylglutamylglutaminylserylleucyllysylisoleucylaspartylthreonylleucylisoleucylglutamylalanylglycylal anylaspartylalanylleucylglutamylleucylglycylisoleucylprolylphenylalanylserylaspartylprolylleucylalanyla spartylglycylprolylthreonylisoleucylglutaminylasparaginylalanylthreonylleucylarginylalanylphenylalanyl alanylalanylglycylvalylthreonylprolylalanylglutaminylcysteinylphenylalanylglutamylmethionylleucylalan ylleucylisoleucylarginylglutaminyllysylhistidylprolylthreonylisoleucylprolylisoleucylglycylleucylleucylmet hionyltyrosylalanylasparaginylleucylvalylphenylalanylasparaginyllysylglycylisoleucylaspartylglutamylp henylalanyltyrosylalanylglutaminylcysteinylglutamyllysylvalylglycylvalylaspartylserylvalylleucylvalylala nylaspartylvalylprolylvalylglutaminylglutamylserylalanylprolylphenylalanylarginylglutaminylalanylalanyl leucylarginylhistidylasparaginylvalylalanylprolylisoleucylphenylalanylisoleucylcysteinylprolylprolylaspa rtylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylgl ycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparagi nylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyros ylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolyl aspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylse rylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylgl utamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylm ethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutamin ylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleuc ylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenylalanylgluta minylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphe nylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylarginylp henylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleucylasp artylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreonylarginylas paraginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylserylprolylthre onylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartylaspartylalany lthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalylasparaginylgl utamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylthreonylphe nylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylalanyltitinmethi onylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylprolylaspartyl alanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycy ltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginyl arginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosyla sparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylasp artylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylg lycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylgluta mylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmeth ionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylp 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hreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartylaspartylalanylthre onylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalylasparaginylglutam ylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylthreonylphenylala nylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylalanyltitinmethionylgl utaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylprolylaspartylalanyl aspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyros ylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginylarginy lalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylaspara ginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylg lutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycyls erylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylpr olylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyll ysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenyl alanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleucyl asparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenylalanylglutaminylthr eonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphenylalan ylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylarginylphenylal anylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleucylaspartylpr 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partylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonylt yrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginylarginylalanylalan ylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanyl alanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylv alyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanyli soleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylprolylglutam yllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanyl alanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenylalanylvaly lphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleucylasparagin 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ylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylalanyltitinmethionylglutaminylarginy ltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylprolylaspartylalanylaspartylaspart ylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonyltyros ylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginylarginylalanylalanylle ucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanylalan ylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalylly sylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisole ucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylprolylglutamyllys ylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanylalan ylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenylalanylvalylphe nylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleucylasparaginylval ylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenylalanylglutaminylthreonylglutamin ylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphenylalanylserylglutamin ylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylarginylphenylalanylprolylglycy laspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleucylaspartylprolylleucylisoleu cylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreonylarginylasparaginylarginylisoleu cylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylserylprolylthreonylthreonylalanylglut amylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartylaspartylalanylthreonylvalylalanyliso leucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalylasparaginylglutamylleucylvalylargin ylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylthreonylphenylalanylglutamylseryl methionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylalanyltitinmethionylglutaminylarginylt yrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylprolylaspartylalanylaspartylaspartyl aspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosyl leucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginylarginylalanylalanylleuc ylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanylalanyl prolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalyllys ylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisoleu 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soleucylglutamylvalylglutamylasparaginylglutaminylglutaminylserylprolylthreonylthreonylalanylglutam ylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartylaspartylalanylthreonylvalylalanylisoleu cylarginylserylalanylasparaginylisoleucylasparaginylleucylvalylasparaginylglutamylleucylvalylarginylgl ycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylthreonylphenylalanylglutamylserylmet hionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylalanyltitinmethionylglutaminylarginyltyros ylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylprolylaspartylalanylaspartylaspartylasp artylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleu cylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginylarginylalanylalanylleucylp rolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanylalanylprol ylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalyllysylal anylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisoleucylv alyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylprolylglutamyllysylmet hionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanylalanylthre onylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenylalanylvalylphenylala nylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleucylasparaginylvalylcyst einylthreonylserylserylleucylglycylasparaginylglutaminylphenylalanylglutaminylthreonylglutaminylgluta minylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphenylalanylserylglutaminylvalyl tryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylarginylphenylalanylprolylglycylaspar 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anylprolylisoleucylphenylalanylisoleucylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleuc ylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucyl serylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginylarginylalanylalanylleucylprolylleuc ylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolyll eucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalan ylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysyli soleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylprolylglutamyllysylmethionylle ucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanylalanylthreonylargi nylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenylalanylvalylphenylalanylleuc ylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthr eonylserylserylleucylglycylasparaginylglutaminylphenylalanylglutaminylthreonylglutaminylglutaminylal anylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptop hyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylarginylphenylalanylprolylglycylaspartylvalyl tyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleucylaspartylprolylleucylisoleucylthreonylal anylleucylleucylglycylthreonylphenylalanylaspartylthreonylarginylasparaginylarginylisoleucylisoleucylg lutamylvalylglutamylasparaginylglutaminylglutaminylserylprolylthreonylthreonylalanylglutamylthreonyll eucylaspartylalanylthreonylarginylarginylvalylaspartylaspartylalanylthreonylvalylalanylisoleucylarginyl serylalanylasparaginylisoleucylasparaginylleucylvalylasparaginylglutamylleucylvalylarginylglycylthreo nylglycylleucyltyrosylasparaginylglutaminylasparaginylthreonylphenylalanylglutamylserylmethionylser ylglycylleucylvalyltryptophylthreonylserylalanylprolylalanyltitinmethionylglutaminylarginyltyrosylglutam ylserylleucylphenylalanylalanylisoleucylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleuc ylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucyl serylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginylarginylalanylalanylleucylprolylleuc ylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolyll eucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalan ylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysyli soleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylprolylglutamyllysylmethionylle 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anylleucylleucylglycylthreonylphenylalanylaspartylthreonylarginylasparaginylarginylisoleucylisoleucylg lutamylvalylglutamylasparaginylglutaminylglutaminylserylprolylthreonylthreonylalanylglutamylthreonyll eucylaspartylalanylthreonylarginylarginylvalylaspartylaspartylalanylthreonylvalylalanylisoleucylarginyl serylalanylasparaginylisoleucylasparaginylleucylvalylasparaginylglutamylleucylvalylarginylglycylthreo nylglycylleucyltyrosylasparaginylglutaminylasparaginylthreonylphenylalanylglutamylserylmethionylser ylglycylleucylvalyltryptophylthreonylserylalanylprolylalanyltitinmethionylglutaminylarginyltyrosylglutam ylserylleucylphenylalanylalanylisoleucylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleuc ylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucyl serylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginylarginylalanylalanylleucylprolylleuc ylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolyll eucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalan ylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysyli soleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylprolylglutamyllysylmethionylle ucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanylalanylthreonylargi nylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenylalanylvalylphenylalanylleuc ylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthr eonylserylserylleucylglycylasparaginylglutaminylphenylalanylglutaminylthreonylglutaminylglutaminylal anylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptop 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rylalanylprolylphenylalanylarginylglutaminylalanylalanylleucylarginylhistidylasparaginylvalylalanylproly lisoleucylphenylalanylisoleucylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylar ginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylargin ylalanylglycylvalylthreonylglycylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparagi nylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylgluta minylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucyl aspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylis oleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanyl alanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetyls 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sparaginylisoleucylasparaginylleucylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleuc yltyrosylasparaginylglutaminylasparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucyl valyltryptophylthreonylserylalanylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylp henylalanylalanylisoleucylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginyl glutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylala nylglycylvalylthreonylglycylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhi stidylleucylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminyl glycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspa rtylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleuc 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anylmethionylthreonylthreonylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylglutaminyll eucyllysylglutamylarginyllysylglutamylglycylalanylphenylalanylvalylprolylphenylalanylvalylthreonylleuc ylglycylaspartylprolylglycylisoleucylglutamylglutaminylserylleucyllysylisoleucylaspartylthreonylleucylis oleucylglutamylalanylglycylalanylaspartylalanylleucylglutamylleucylglycylisoleucylprolylphenylalanylse rylaspartylprolylleucylalanylaspartylglycylprolylthreonylisoleucylglutaminylasparaginylalanylthreonylle ucylarginylalanylphenylalanylalanylalanylglycylvalylthreonylprolylalanylglutaminylcysteinylphenylalany lglutamylmethionylleucylalanylleucylisoleucylarginylglutaminyllysylhistidylprolylthreonylisoleucylprolyli soleucylglycylleucylleucylmethionyltyrosylalanylasparaginylleucylvalylphenylalanylasparaginyllysylgly cylisoleucylaspartylglutamylphenylalanyltyrosylalanylglutaminylcysteinylglutamyllysylvalylglycylvalyla 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anylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhi stidylasparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenyl alanylvalylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthre onylserylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspart ylprolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginyl glutaminylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylv alylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutami nylserylthreonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylas paraginylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylal anylaspartylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylgluta minylglutaminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylar ginylvalylaspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylaspar aginylleucylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylgluta minylasparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylse rylalanylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleuc ylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalany lseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylgly cylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanylly 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utamylphenylalanyltyrosylalanylglutaminylcysteinylglutamyllysylvalylglycylvalylaspartylserylvalylleucyl valylalanylaspartylvalylprolylvalylglutaminylglutamylserylalanylprolylphenylalanylarginylglutaminylalan ylalanylleucylarginylhistidylasparaginylvalylalanylprolylisoleucylphenylalanylisoleucylcysteinylprolylpr olylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycyl arginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse 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lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla spartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalyla sparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylt hreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylala nyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylp rolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycy larginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu 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rolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycy larginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla spartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalyla sparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylt hreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylala nyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylmethionylthreonylthreon ylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylglutaminylleucyllysylglutamylarginyllys ylglutamylglycylalanylphenylalanylvalylprolylphenylalanylvalylthreonylleucylglycylaspartylprolylglycylis oleucylglutamylglutaminylserylleucyllysylisoleucylaspartylthreonylleucylisoleucylglutamylalanylglycylal anylaspartylalanylleucylglutamylleucylglycylisoleucylprolylphenylalanylserylaspartylprolylleucylalanyla spartylglycylprolylthreonylisoleucylglutaminylasparaginylalanylthreonylleucylarginylalanylphenylalanyl alanylalanylglycylvalylthreonylprolylalanylglutaminylcysteinylphenylalanylglutamylmethionylleucylalan ylleucylisoleucylarginylglutaminyllysylhistidylprolylthreonylisoleucylprolylisoleucylglycylleucylleucylmet 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rylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylgl utamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylm ethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutamin ylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleuc ylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenylalanylgluta minylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphe nylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylarginylp henylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleucylasp artylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreonylarginylas paraginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylserylprolylthre onylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartylaspartylalany lthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalylasparaginylgl utamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylthreonylphe nylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylalanyltitinmethi onylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylprolylaspartyl alanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycy ltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginyl arginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosyla sparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylasp artylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylg lycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylgluta mylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmeth ionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylp henylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucyll eucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenylalanylglutami nylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylpheny lalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylarginylphe 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hreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphenylala nylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylarginylphenyl alanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleucylaspartyl prolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreonylarginylaspara ginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylserylprolylthreonylt hreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartylaspartylalanylthre onylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalylasparaginylglutam ylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylthreonylphenylala nylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylalanyltitinmethionylgl utaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylprolylaspartylalanyl aspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyros ylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginylarginy lalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylaspara ginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylg lutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycyls erylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylpr olylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyll ysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenyl alanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleucyl asparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenylalanylglutaminylth reonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphenylalan ylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylarginylphenylal anylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleucylaspartylpr olylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreonylarginylasparagi nylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylserylprolylthreonylthr eonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartylaspartylalanylthreon ylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalylasparaginylglutamyll eucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylthreonylphenylalan ylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylalanyltitinmethionylglu taminylarginyltyrosylglutamylserylleucylphenylalanylalanylmethionylthreonylthreonylglutaminylarginylt yrosylglutamylserylleucylphenylalanylalanylglutaminylleucyllysylglutamylarginyllysylglutamylglycylala nylphenylalanylvalylprolylphenylalanylvalylthreonylleucylglycylaspartylprolylglycylisoleucylglutamylglu taminylserylleucyllysylisoleucylaspartylthreonylleucylisoleucylglutamylalanylglycylalanylaspartylalanyll eucylglutamylleucylglycylisoleucylprolylphenylalanylserylaspartylprolylleucylalanylaspartylglycylprolylt hreonylisoleucylglutaminylasparaginylalanylthreonylleucylarginylalanylphenylalanylalanylalanylglycylv alylthreonylprolylalanylglutaminylcysteinylphenylalanylglutamylmethionylleucylalanylleucylisoleucylar ginylglutaminyllysylhistidylprolylthreonylisoleucylprolylisoleucylglycylleucylleucylmethionyltyrosylalany lasparaginylleucylvalylphenylalanylasparaginyllysylglycylisoleucylaspartylglutamylphenylalanyltyrosyl alanylglutaminylcysteinylglutamyllysylvalylglycylvalylaspartylserylvalylleucylvalylalanylaspartylvalylpro lylvalylglutaminylglutamylserylalanylprolylphenylalanylarginylglutaminylalanylalanylleucylarginylhistidy lasparaginylvalylalanylprolylisoleucylphenylalanylisoleucylcysteinylprolylprolylaspartylalanylaspartylas partylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonylt yrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginylarginylalanylalan ylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanyl alanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylv alyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanyli soleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylprolylglutam yllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanyl alanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenylalanylvaly lphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleucylasparagin ylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenylalanylglutaminylthreonylgluta minylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphenylalanylserylgluta minylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylarginylphenylalanylprolylgl ycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleucylaspartylprolylleucyliso 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ucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanylalan ylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalylly sylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisole ucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylprolylglutamyllys ylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanylalan ylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenylalanylvalylphe nylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleucylasparaginylval ylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenylalanylglutaminylthreonylglutamin ylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphenylalanylserylglutamin 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aspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosyl leucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginylarginylalanylalanylleuc ylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanylalanyl prolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalyllys ylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisoleu cylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylglutamylprolylglutamyllysyl methionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanylalanylt hreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenylalanylvalylpheny lalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleucylasparaginylvalylc 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ylisoleucylasparaginylleucylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosyl asparaginylglutaminylasparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryp tophylthreonylserylalanylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylala nylalanylisoleucylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutamin ylisoleucylalanylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycyl valylthreonylglycylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleu cylvalylalanyllysylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylph enylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalany lglycylalanylalanylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutam ylglutaminylhistidylasparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllys ylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylseryl isoleucylthreonylserylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophyl alanylaspartylprolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycy lasparaginylglutaminylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreon ylglutaminylvalylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanyl prolylglutaminylserylthreonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylargi nyltyrosylasparaginylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreo nylphenylalanylaspartylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylaspar 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rylaspartylprolylleucylalanylaspartylglycylprolylthreonylisoleucylglutaminylasparaginylalanylthreonylle ucylarginylalanylphenylalanylalanylalanylglycylvalylthreonylprolylalanylglutaminylcysteinylphenylalany lglutamylmethionylleucylalanylleucylisoleucylarginylglutaminyllysylhistidylprolylthreonylisoleucylprolyli soleucylglycylleucylleucylmethionyltyrosylalanylasparaginylleucylvalylphenylalanylasparaginyllysylgly cylisoleucylaspartylglutamylphenylalanyltyrosylalanylglutaminylcysteinylglutamyllysylvalylglycylvalyla spartylserylvalylleucylvalylalanylaspartylvalylprolylvalylglutaminylglutamylserylalanylprolylphenylalany larginylglutaminylalanylalanylleucylarginylhistidylasparaginylvalylalanylprolylisoleucylphenylalanylisol eucylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylal anylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreony lglycylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalan yllysylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylgl ycylisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalany lalanylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminyl histidylasparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphen ylalanylvalylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylth reonylserylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspa rtylprolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparagin ylglutaminylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminy lvalylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylgluta minylserylthreonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosyl asparaginylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenyl alanylaspartylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglut aminylglutaminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginyla rginylvalylaspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylaspa raginylleucylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglut aminylasparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonyls erylalanylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleu cylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalan ylseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylgl ycylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyll ysylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglyc ylisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylal anylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhi stidylasparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenyl alanylvalylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthre onylserylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspart ylprolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginyl glutaminylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylv alylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutami nylserylthreonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylas paraginylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylal anylaspartylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylgluta minylglutaminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylar ginylvalylaspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylaspar aginylleucylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylgluta minylasparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylse rylalanylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleuc ylcysteinylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalany lseryltyrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylgly cylalanylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanylly sylleucyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycy lisoleucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylala nylglycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhist idylasparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylal anylvalylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreo nylserylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartyl prolylisoleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylgl utaminylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylval ylglutaminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminy lserylthreonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylaspar aginylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalany laspartylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminy lglutaminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylv alylaspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginyl leucylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminyl asparaginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylala nylprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcyst einylprolylprolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylserylt yrosylglycylarginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylala nylglutamylasparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleu cyllysylglutamyltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisole ucylserylalanylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylgl ycylalanylisoleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidyla sparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylv alylglutaminylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylser ylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolyli soleucylglutamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutami nylphenylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglut aminylglutaminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylseryl threonylvalylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparagin ylalanylvalylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylasp artylthreonylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglut aminylserylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalyl aspartylaspartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleu cylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasp araginylthreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanyl prolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylmethionylthreon ylthreonylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylglutaminylleucyllysylglutamylar ginyllysylglutamylglycylalanylphenylalanylvalylprolylphenylalanylvalylthreonylleucylglycylaspartylprolyl glycylisoleucylglutamylglutaminylserylleucyllysylisoleucylaspartylthreonylleucylisoleucylglutamylalanyl glycylalanylaspartylalanylleucylglutamylleucylglycylisoleucylprolylphenylalanylserylaspartylprolylleucy lalanylaspartylglycylprolylthreonylisoleucylglutaminylasparaginylalanylthreonylleucylarginylalanylphen ylalanylalanylalanylglycylvalylthreonylprolylalanylglutaminylcysteinylphenylalanylglutamylmethionylleu cylalanylleucylisoleucylarginylglutaminyllysylhistidylprolylthreonylisoleucylprolylisoleucylglycylleucylle ucylmethionyltyrosylalanylasparaginylleucylvalylphenylalanylasparaginyllysylglycylisoleucylaspartylgl utamylphenylalanyltyrosylalanylglutaminylcysteinylglutamyllysylvalylglycylvalylaspartylserylvalylleucyl valylalanylaspartylvalylprolylvalylglutaminylglutamylserylalanylprolylphenylalanylarginylglutaminylalan ylalanylleucylarginylhistidylasparaginylvalylalanylprolylisoleucylphenylalanylisoleucylcysteinylprolylpr olylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycyl arginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla spartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalyla sparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylt hreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylala nyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylp rolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycy larginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla spartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalyla sparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylt hreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylala nyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylp rolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycy larginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva lylleucylaspartylprolylleucylisoleucylthreonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreo nylarginylasparaginylarginylisoleucylisoleucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthreonylalanylglutamylthreonylleucylaspartylalanylthreonylarginylarginylvalylaspartyla spartylalanylthreonylvalylalanylisoleucylarginylserylalanylasparaginylisoleucylasparaginylleucylvalyla sparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasparaginylt hreonylphenylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylala nyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylp rolylaspartylalanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycy larginylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamyl asparaginylarginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylgluta myltyrosylasparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylala nylprolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylis oleucylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginyli soleucylglutamylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutami nylprolylmethionyllysylalanylalanylthreonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylser ylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylgl utamylleucylleucylasparaginylvalylcysteinylthreonylserylserylleucylglycylasparaginylglutaminylphenyl alanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglu taminylphenylalanylserylglutaminylvalyltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylv alylarginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylva 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nylalanylglutamylserylmethionylserylglycylleucylvalyltryptophylthreonylserylalanylprolylalanyltitinmethi onylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcysteinylprolylprolylaspartyl alanylaspartylaspartylaspartylleucylleucylarginylglutaminylisoleucylalanylseryltyrosylglycylarginylglycy ltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylasparaginyl arginylalanylalanylleucylprolylleucylasparaginylhistidylleucylvalylalanyllysylleucyllysylglutamyltyrosyla sparaginylalanylalanylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylasp artylglutaminylvalyllysylalanylalanylisoleucylaspartylalanylglycylalanylalanylglycylalanylisoleucylserylg lycylserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylasparaginylisoleucylgluta mylprolylglutamyllysylmethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmeth 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ethionylleucylalanylalanylleucyllysylvalylphenylalanylvalylglutaminylprolylmethionyllysylalanylalanylthr eonylarginylacetylseryltyrosylserylisoleucylthreonylserylprolylserylglutaminylphenylalanylvalylphenylal anylleucylserylserylvalyltryptophylalanylaspartylprolylisoleucylglutamylleucylleucylasparaginylvalylcys teinylthreonylserylserylleucylglycylasparaginylglutaminylphenylalanylglutaminylthreonylglutaminylglut aminylalanylarginylthreonylthreonylglutaminylvalylglutaminylglutaminylphenylalanylserylglutaminylval yltryptophyllysylprolylphenylalanylprolylglutaminylserylthreonylvalylarginylphenylalanylprolylglycylasp artylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleucylaspartylprolylleucylisoleucylth reonylalanylleucylleucylglycylthreonylphenylalanylaspartylthreonylarginylasparaginylarginylisoleucylis oleucylglutamylvalylglutamylasparaginylglutaminylglutaminylserylglutamylasparaginylglutaminylgluta minylserylglutamylasparaginylglutaminylglutaminylserxisoleucine

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by: rsylestine

Posted: Dec 7, 2012 / 07:00 PM CST

Updated: Feb 4, 2021 / 10:59 AM CST

This is an archived article and the information in the article may be outdated. Please look at the time stamp on the story to see when it was last updated.

ESQUIRE, RUSSIA (CW39) – If someone says they’d like to have a word with you, make sure it’s not this one! It’s the longest word in English and if you want to read it, here’s a look. A word of warning… the “word” takes about 3.5 hours to say.

The word is 189,819 letters long. It’s actually the name of a giant protein called Titin. Proteins are usually named by mashing-up the names of the chemicals making them. And since Titin is the largest protein ever discovered, its name had to be equally as large.

If you’re interested in something ‘shorter,’ here are several more $10 words to wrap your mind around.

1. Pneumonoultramicroscopicsilicovolcanoconiosis (45 letters)

Pneumoconiosis comes after you inhale very fine silicate or quartz dust.

2. Hippopotomonstrosesquippedaliophobia (36 letters)

Have a fear of long words? That’s called: Hippopotomonstrosesquippedaliophobia and it’s one of the longest words in the dictionary.

3. Supercalifragilisticexpialidocious (34 letters)

Perhaps the most fun word of all, is the word made popular by the film Mary Poppins. Supercalifragilisticexpialidocious.

4. Pseudopseudohypoparathyroidism (30 letters)

Pseudohypoparathyroidism is characterized by normal levels of calcium and phosphorus in the blood.

5. Floccinaucinihilipilification (29 letters)

The longest unchallenged nontechnical word that not all directories recognize, is floccinaucinihilipilification – the act or habit of describing or regarding something as unimportant, of having no value or being worthless. Often times, it is used in good humor.

Here are more of the longest words out there and how to say them!

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